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n-3 polyunsaturated fatty acids suppress phosphatidylinositol-(4,5)-bisphosphate dependent actin remodeling during CD4+ T cell activation.
MedLine Citation:
PMID:  22250985     Owner:  NLM     Status:  Publisher    
Abstract/OtherAbstract:
n-3 polyunsaturated fatty acids (PUFA), i.e. docosahexaenoic acid (DHA), found in fish oil, exhibit anti-inflammatory properties; however, the molecular mechanisms remain unclear. Since phosphatidylinositol-(4,5)-bisphosphate (PI(4,5)P2) resides in raft domains and DHA can alter the size of rafts, we hypothesized that PI(4,5)P2 and downstream actin remodeling are perturbed by the incorporation of n-3 PUFA into membranes, resulting in suppressed T cell activation. CD4+ T cells isolated from Fat-1 transgenic mice (membranes enriched in n-3 PUFA) exhibited a 50% decrease in PI(4,5)P2. Upon activation by plate bound anti-CD3/anti-CD28 or PMA/ionomycin, Fat-1 CD4+ T cells failed to metabolize PI(4,5)P2. Furthermore, actin remodeling, failed to initiate in Fat-1 CD4+ T cells upon stimulation; however, the defect was reversed by incubation with exogenous PI(4,5)P2. When Fat-1 CD4+ T cells were stimulated with anti-CD3/anti-CD28 coated beads, the Wiskott-Aldrich syndrome protein (WASP) failed to translocate to the immunological synapse. The suppressive phenotype, consisting of defects in PI(4,5)P2 metabolism and actin remodeling, were recapitulated in CD4+ T cells isolated from mice fed a 4% DHA triglyceride-enriched diet. Collectively, these data demonstrate that DHA alters PI(4,5)P2 in CD4+ T cells, thereby suppressing the recruitment of WASP to the IS, and impairing actin remodeling in CD4+ T cells.
Authors:
Tim Y Hou; Jennifer M Monk; Yang-Yi Fan; Rola Barhoumi; Yong Q Chen; Gonzalo M Rivera; David N McMurray; Robert S Chapkin
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Publication Detail:
Type:  JOURNAL ARTICLE     Date:  2012-1-18
Journal Detail:
Title:  The Biochemical journal     Volume:  -     ISSN:  1470-8728     ISO Abbreviation:  -     Publication Date:  2012 Jan 
Date Detail:
Created Date:  2012-1-18     Completed Date:  -     Revised Date:  -    
Medline Journal Info:
Nlm Unique ID:  2984726R     Medline TA:  Biochem J     Country:  -    
Other Details:
Languages:  ENG     Pagination:  -     Citation Subset:  -    
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