Document Detail


A molecularly identified P2Y receptor simultaneously activates phospholipase C and inhibits adenylyl cyclase and is nonselectively activated by all nucleoside triphosphates.
MedLine Citation:
PMID:  10727529     Owner:  NLM     Status:  MEDLINE    
Abstract/OtherAbstract:
We recently cloned and expressed a novel P2Y receptor (tp2y receptor) from a turkey cDNA library. Expression of this receptor in 1321N1 human astrocytoma cells confers nucleotide-dependent stimulation of phospholipase C activity; however, as we demonstrate here, it also confers nucleotide-dependent inhibition of adenylyl cyclase. Both the phospholipase C and adenylyl cyclase responses were promoted by receptor agonists over a similar range of concentrations. Moreover, not only did UTP and ATP activate the avian receptor but ITP, GTP, xanthosine 5'-triphosphate, and CTP were also agonists, with EC(50) values ranging between 0.1 and 1 microM. Similar potencies, rank-order, and selectivity of nucleotide agonists were also demonstrated for intracellular Ca(2+) mobilization measured during a 30-s stimulation under constant superfusion conditions. This observation indicates that receptor activation by nucleoside 5'-triphosphates is not produced by interconversion of these nucleotides into ATP or UTP. Pretreatment of cells with pertussis toxin completely abolished the inhibitory effect of nucleotide agonists on adenylyl cyclase, whereas the activation of phospholipase C was only partially inhibited. These results demonstrate that the avian P2Y receptor is a nucleoside triphosphate receptor of broad agonist selectivity that interacts with both pertussis toxin-insensitive and -sensitive G proteins to activate phospholipase C and to inhibit adenylyl cyclase. This is the first cloned P2Y receptor that is clearly Gi/adenylyl cyclase-linked.
Authors:
J L Boyer; S M Delaney; D Villanueva; T K Harden
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Publication Detail:
Type:  Journal Article; Research Support, Non-U.S. Gov't; Research Support, U.S. Gov't, P.H.S.    
Journal Detail:
Title:  Molecular pharmacology     Volume:  57     ISSN:  0026-895X     ISO Abbreviation:  Mol. Pharmacol.     Publication Date:  2000 Apr 
Date Detail:
Created Date:  2000-05-04     Completed Date:  2000-05-04     Revised Date:  2010-01-14    
Medline Journal Info:
Nlm Unique ID:  0035623     Medline TA:  Mol Pharmacol     Country:  UNITED STATES    
Other Details:
Languages:  eng     Pagination:  805-10     Citation Subset:  IM    
Affiliation:
Department of Pharmacology, The School of Medicine, University of North Carolina, Chapel Hill, North Carolina 7599-7365, USA. boyerl@med.unc.edu
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MeSH Terms
Descriptor/Qualifier:
Adenosine Triphosphate / metabolism
Adenylate Cyclase / antagonists & inhibitors,  metabolism*
Animals
Cloning, Molecular
Cytidine Triphosphate / metabolism
Enzyme Activation
Enzyme Inhibitors
GTP-Binding Proteins / metabolism
Guanosine Triphosphate / metabolism
Humans
Nucleotides / metabolism*
Receptors, Purinergic P2 / genetics,  metabolism*
Tumor Cells, Cultured
Turkeys
Type C Phospholipases / metabolism*
Uridine Triphosphate / metabolism
Grant Support
ID/Acronym/Agency:
GM38213/GM/NIGMS NIH HHS; HL54889/HL/NHLBI NIH HHS
Chemical
Reg. No./Substance:
0/Enzyme Inhibitors; 0/Nucleotides; 0/Receptors, Purinergic P2; 0/purinergic receptor P2Y2; 56-65-5/Adenosine Triphosphate; 63-39-8/Uridine Triphosphate; 65-47-4/Cytidine Triphosphate; 86-01-1/Guanosine Triphosphate; EC 3.1.4.-/Type C Phospholipases; EC 3.6.1.-/GTP-Binding Proteins; EC 4.6.1.1/Adenylate Cyclase

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