Document Detail


mTORC1-mediated cell proliferation, but not cell growth, controlled by the 4E-BPs.
MedLine Citation:
PMID:  20508131     Owner:  NLM     Status:  MEDLINE    
Abstract/OtherAbstract:
The mammalian target of rapamycin complex 1 (mTORC1) integrates mitogen and nutrient signals to control cell proliferation and cell size. Hence, mTORC1 is implicated in a large number of human diseases--including diabetes, obesity, heart disease, and cancer--that are characterized by aberrant cell growth and proliferation. Although eukaryotic translation initiation factor 4E-binding proteins (4E-BPs) are critical mediators of mTORC1 function, their precise contribution to mTORC1 signaling and the mechanisms by which they mediate mTORC1 function have remained unclear. We inhibited the mTORC1 pathway in cells lacking 4E-BPs and analyzed the effects on cell size, cell proliferation, and cell cycle progression. Although the 4E-BPs had no effect on cell size, they inhibited cell proliferation by selectively inhibiting the translation of messenger RNAs that encode proliferation-promoting proteins and proteins involved in cell cycle progression. Thus, control of cell size and cell cycle progression appear to be independent in mammalian cells, whereas in lower eukaryotes, 4E-BPs influence both cell growth and proliferation.
Authors:
Ryan J O Dowling; Ivan Topisirovic; Tommy Alain; Michael Bidinosti; Bruno D Fonseca; Emmanuel Petroulakis; Xiaoshan Wang; Ola Larsson; Anand Selvaraj; Yi Liu; Sara C Kozma; George Thomas; Nahum Sonenberg
Publication Detail:
Type:  Journal Article; Research Support, N.I.H., Extramural; Research Support, Non-U.S. Gov't; Research Support, U.S. Gov't, Non-P.H.S.    
Journal Detail:
Title:  Science (New York, N.Y.)     Volume:  328     ISSN:  1095-9203     ISO Abbreviation:  Science     Publication Date:  2010 May 
Date Detail:
Created Date:  2010-05-28     Completed Date:  2010-06-08     Revised Date:  2011-09-26    
Medline Journal Info:
Nlm Unique ID:  0404511     Medline TA:  Science     Country:  United States    
Other Details:
Languages:  eng     Pagination:  1172-6     Citation Subset:  IM    
Affiliation:
Department of Biochemistry and Goodman Cancer Research Centre, McGill University, Montreal, Quebec H3A 1A3, Canada.
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MeSH Terms
Descriptor/Qualifier:
Animals
Carrier Proteins / genetics,  metabolism*
Cell Cycle
Cell Enlargement*
Cell Line
Cell Proliferation*
Cell Size
Cell Survival
Eukaryotic Initiation Factors / genetics,  metabolism*
Humans
Mice
Mice, Knockout
Phosphoproteins / genetics,  metabolism*
Phosphorylation
Protein Biosynthesis
RNA, Messenger / genetics,  metabolism
Ribosomal Protein S6 Kinases / metabolism
Signal Transduction
Sirolimus / pharmacology
Transcription Factors / metabolism*
Grant Support
ID/Acronym/Agency:
P50 NS057531/NS/NINDS NIH HHS; P50 NS057531-01A2/NS/NINDS NIH HHS; R01 DK73802/DK/NIDDK NIH HHS; U01 CA84292-06/CA/NCI NIH HHS; //Canadian Institutes of Health Research; //Howard Hughes Medical Institute
Chemical
Reg. No./Substance:
0/Carrier Proteins; 0/Eif4ebp1 protein, mouse; 0/Eif4ebp2 protein, mouse; 0/Eukaryotic Initiation Factors; 0/Phosphoproteins; 0/RNA, Messenger; 0/Transcription Factors; 0/mTORC1 complex, human; 0/mTORC1 complex, mouse; 53123-88-9/Sirolimus; EC 2.7.11.1/Ribosomal Protein S6 Kinases
Comments/Corrections

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