Document Detail


The lipid A from Vibrio fischeri lipopolysaccharide: a unique structure bearing a phosphoglycerol moiety.
MedLine Citation:
PMID:  21498521     Owner:  NLM     Status:  MEDLINE    
Abstract/OtherAbstract:
Vibrio fischeri, a bioluminescent marine bacterium, exists in an exclusive symbiotic relationship with the Hawaiian bobtail squid, Euprymna scolopes, whose light organ it colonizes. Previously, it has been shown that the lipopolysaccharide (LPS) or free lipid A of V. fischeri can trigger morphological changes in the juvenile squid's light organ that occur upon colonization. To investigate the structural features that might be responsible for this phenomenon, the lipid A from V. fischeri ES114 LPS was isolated and characterized by multistage mass spectrometry (MS(n)). A microheterogeneous mixture of mono- and diphosphorylated diglucosamine disaccharides was observed with variable states of acylation ranging from tetra- to octaacylated forms. All lipid A species, however, contained a set of conserved primary acyl chains consisting of an N-linked C14:0(3-OH) at the 2-position, an unusual N-linked C14:1(3-OH) at the 2'-position, and two O-linked C12:0(3-OH) fatty acids at the 3- and 3'-positions. The fatty acids found in secondary acylation were considerably more variable, with either a C12:0 or C16:1 at the 2-position, C14:0 or C14:0(3-OH) at the 2'-position, and C12:0 or no substituent at the 3'-position. Most surprising was the presence of an unusual set of modifications at the secondary acylation site of the 3-position consisting of phosphoglycerol (GroP), lysophosphatidic acid (GroP bearing C12:0, C16:0, or C16:1), or phosphatidic acid (GroP bearing either C16:0 + C12:0 or C16:0 + C16:1). Given their unusual nature, it is possible that these features of the V. fischeri lipid A may underlie the ability of E. scolopes to recognize its symbiotic partner.
Authors:
Nancy J Phillips; Dawn M Adin; Eric V Stabb; Margaret J McFall-Ngai; Michael A Apicella; Bradford W Gibson
Publication Detail:
Type:  Journal Article; Research Support, N.I.H., Extramural; Research Support, Non-U.S. Gov't     Date:  2011-04-15
Journal Detail:
Title:  The Journal of biological chemistry     Volume:  286     ISSN:  1083-351X     ISO Abbreviation:  J. Biol. Chem.     Publication Date:  2011 Jun 
Date Detail:
Created Date:  2011-06-13     Completed Date:  2011-08-30     Revised Date:  2012-01-09    
Medline Journal Info:
Nlm Unique ID:  2985121R     Medline TA:  J Biol Chem     Country:  United States    
Other Details:
Languages:  eng     Pagination:  21203-19     Citation Subset:  IM    
Affiliation:
Department of Pharmaceutical Chemistry, University of California, San Francisco, California 94143, USA.
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MeSH Terms
Descriptor/Qualifier:
Aliivibrio fischeri / metabolism*
Animals
Decapodiformes / microbiology*
Fatty Acids / chemistry
Gas Chromatography-Mass Spectrometry / methods
Glycerophosphates / chemistry
Lipid A / chemistry*
Lipopolysaccharides / metabolism*
Models, Chemical
Phosphatidic Acids / chemistry
Phosphorylation
Spectrometry, Mass, Electrospray Ionization / methods
Spectrometry, Mass, Matrix-Assisted Laser Desorption-Ionization
Symbiosis
Tandem Mass Spectrometry / methods
Grant Support
ID/Acronym/Agency:
AI-50661/AI/NIAID NIH HHS; P30 CA082103/CA/NCI NIH HHS; R01 AI024616-23/AI/NIAID NIH HHS
Chemical
Reg. No./Substance:
0/Fatty Acids; 0/Glycerophosphates; 0/Lipid A; 0/Lipopolysaccharides; 0/Phosphatidic Acids; 57-03-4/alpha-glycerophosphoric acid

From MEDLINE®/PubMed®, a database of the U.S. National Library of Medicine


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