Document Detail


The Mus81/Mms4 endonuclease acts independently of double-Holliday junction resolution to promote a distinct subset of crossovers during meiosis in budding yeast.
MedLine Citation:
PMID:  12750322     Owner:  NLM     Status:  MEDLINE    
Abstract/OtherAbstract:
Current models for meiotic recombination require that crossovers derive from the resolution of a double-Holliday junction (dHJ) intermediate. In prokaryotes, enzymes responsible for HJ resolution are well characterized but the identification of a eukaryotic nuclear HJ resolvase has been elusive. Indirect evidence suggests that MUS81 from humans and fission yeast encodes a HJ resolvase. We provide three lines of evidence that Mus81/Mms4 is not the major meiotic HJ resolvase in S. cerevisiae: (1) MUS81/MMS4 is required to form only a distinct subset of crossovers; (2) rather than accumulating, dHJ intermediates are reduced in an mms4 mutant; and (3) expression of a bacterial HJ resolvase has no suppressive effect on mus81 meiotic phenotypes. Our analysis also reveals the existence of two distinct classes of crossovers in budding yeast. Class I is dependent upon MSH4/MSH5 and exhibits crossover interference, while class II is dependent upon MUS81/MMS4 and exhibits no interference. mms4 specifically reduces crossing over on small chromosomes, which are known to undergo less interference. The correlation between recombination rate and degree of interference to chromosome size may therefore be achieved by modulating the balance between class I/class II crossovers.
Authors:
Teresa de los Santos; Neil Hunter; Cindy Lee; Brittany Larkin; Josef Loidl; Nancy M Hollingsworth
Publication Detail:
Type:  Journal Article; Research Support, Non-U.S. Gov't; Research Support, U.S. Gov't, P.H.S.    
Journal Detail:
Title:  Genetics     Volume:  164     ISSN:  0016-6731     ISO Abbreviation:  Genetics     Publication Date:  2003 May 
Date Detail:
Created Date:  2003-05-16     Completed Date:  2004-01-30     Revised Date:  2013-02-19    
Medline Journal Info:
Nlm Unique ID:  0374636     Medline TA:  Genetics     Country:  United States    
Other Details:
Languages:  eng     Pagination:  81-94     Citation Subset:  IM    
Affiliation:
Institute for Cell and Developmental Biology, Biochemistry and Cell Biology, State University of New York, Stony Brook, New York 11794-5215, USA.
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MeSH Terms
Descriptor/Qualifier:
Crossing Over, Genetic*
DNA, Cruciform
DNA-Binding Proteins / genetics,  metabolism*
Endonucleases*
Flap Endonucleases
Meiosis / genetics,  physiology
Saccharomyces cerevisiae / enzymology,  genetics*
Saccharomyces cerevisiae Proteins*
Trans-Activators / genetics,  metabolism*
Grant Support
ID/Acronym/Agency:
GM-50717/GM/NIGMS NIH HHS
Chemical
Reg. No./Substance:
0/DNA, Cruciform; 0/DNA-Binding Proteins; 0/Saccharomyces cerevisiae Proteins; 0/Trans-Activators; EC 3.1.-/Endonucleases; EC 3.1.-/Flap Endonucleases; EC 3.1.-/MUS81 protein, S cerevisiae; EC 3.1.22.-/MMS4 protein, S cerevisiae
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