Document Detail


A cellular timetable of autumn senescence.
MedLine Citation:
PMID:  16299183     Owner:  NLM     Status:  MEDLINE    
Abstract/OtherAbstract:
We have studied autumn leaf senescence in a free-growing aspen (Populus tremula) by following changes in pigment, metabolite and nutrient content, photosynthesis, and cell and organelle integrity. The senescence process started on September 11, 2003, apparently initiated solely by the photoperiod, and progressed steadily without any obvious influence of other environmental signals. For example, after this date, senescing leaves accumulated anthocyanins in response to conditions inducing photooxidative stress, but at the beginning of September the leaves did not. Degradation of leaf constituents took place over an 18-d period, and, although the cells in each leaf did not all senesce in parallel, senescence in the tree as a whole was synchronous. Lutein and beta-carotene were degraded in parallel with chlorophyll, whereas neoxanthin and the xanthophyll cycle pigments were retained longer. Chloroplasts in each cell were rapidly converted to gerontoplasts and many, although not all, cells died. From September 19, when chlorophyll levels had dropped by 50%, mitochondrial respiration provided the energy for nutrient remobilization. Remobilization seemed to stop on September 29, probably due to the cessation of phloem transport, but, up to abscission of the last leaves (over 1 week later), some cells were metabolically active and had chlorophyll-containing gerontoplasts. About 80% of the nitrogen and phosphorus was remobilized, and on September 29 a sudden change occurred in the delta15N of the cellular content, indicating that volatile compounds may have been released.
Authors:
Johanna Keskitalo; Gustaf Bergquist; Per Gardeström; Stefan Jansson
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Publication Detail:
Type:  Journal Article; Research Support, Non-U.S. Gov't     Date:  2005-11-18
Journal Detail:
Title:  Plant physiology     Volume:  139     ISSN:  0032-0889     ISO Abbreviation:  Plant Physiol.     Publication Date:  2005 Dec 
Date Detail:
Created Date:  2005-12-12     Completed Date:  2006-03-20     Revised Date:  2010-09-21    
Medline Journal Info:
Nlm Unique ID:  0401224     Medline TA:  Plant Physiol     Country:  United States    
Other Details:
Languages:  eng     Pagination:  1635-48     Citation Subset:  IM    
Affiliation:
Umeå Plant Science Centre, Department of Plant Physiology, Umeå University, S-901 87 Umea, Sweden. johanna.keskitalo@plantphys.umu.se
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MeSH Terms
Descriptor/Qualifier:
Anthocyanins / metabolism
Carotenoids / metabolism
Chlorophyll / metabolism
Chloroplasts / metabolism
Microscopy, Electron
Mitochondria / metabolism
Nitrogen / metabolism
Phosphorus / metabolism
Photobiology
Photoperiod
Photosynthesis
Pigments, Biological / metabolism
Plant Leaves / cytology,  growth & development*,  metabolism
Populus / cytology,  growth & development*,  metabolism
Seasons*
Chemical
Reg. No./Substance:
0/Anthocyanins; 0/Pigments, Biological; 1406-65-1/Chlorophyll; 36-88-4/Carotenoids; 7723-14-0/Phosphorus; 7727-37-9/Nitrogen
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