Document Detail


Zyg-11 and cul-2 regulate progression through meiosis II and polarity establishment in C. elegans.
MedLine Citation:
PMID:  15215208     Owner:  NLM     Status:  MEDLINE    
Abstract/OtherAbstract:
The mechanisms that ensure coupling between meiotic cell cycle progression and subsequent developmental events, including specification of embryonic axes, are poorly understood. Here, we establish that zyg-11 and the cullin cul-2 promote the metaphase-to-anaphase transition and M phase exit at meiosis II in Caenorhabditis elegans. Our results indicate that ZYG-11 acts with a CUL-2-based E3 ligase that is essential at meiosis II and that functions redundantly with the anaphase-promoting complex/cyclosome at meiosis I. Our data also indicate that delayed M phase exit in zyg-11(RNAi) embryos is due to accumulation of the B type cyclin CYB-3. We demonstrate that PAR proteins and P granules become polarized in an inverted manner during the meiosis II delay resulting from zyg-11 or cul-2 inactivation, and that zyg-11 and cul-2 can regulate polarity establishment independently of a role in cell cycle progression. Furthermore, we find that microtubules appear dispensable for ectopic polarity during the meiosis II delay in zyg-11(RNAi) embryos, as well as for AP polarity during the first mitotic cell cycle in wild-type embryos. Our findings suggest a model in which a CUL-2-based E3 ligase promotes cell cycle progression and prevents polarity establishment during meiosis II, and in which the centrosome acts as a cue to polarize the embryo along the AP axis after exit from the meiotic cell cycle.
Authors:
Rémi Sonneville; Pierre Gönczy
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Publication Detail:
Type:  Journal Article; Research Support, Non-U.S. Gov't     Date:  2004-06-23
Journal Detail:
Title:  Development (Cambridge, England)     Volume:  131     ISSN:  0950-1991     ISO Abbreviation:  Development     Publication Date:  2004 Aug 
Date Detail:
Created Date:  2004-07-20     Completed Date:  2004-09-30     Revised Date:  2009-11-19    
Medline Journal Info:
Nlm Unique ID:  8701744     Medline TA:  Development     Country:  England    
Other Details:
Languages:  eng     Pagination:  3527-43     Citation Subset:  IM    
Affiliation:
ISREC (Swiss Institute for Experimental Cancer Research Boveresses, CH-1066 Epalinges/Lausanne, Switzerland.
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MeSH Terms
Descriptor/Qualifier:
Animals
Biological Markers
Body Patterning*
Caenorhabditis elegans / embryology*
Caenorhabditis elegans Proteins / genetics,  metabolism*
Cell Cycle Proteins / genetics,  metabolism*
Cell Polarity
Cullin Proteins / genetics,  metabolism*
Cyclin B / genetics,  metabolism*
Cyclin B1
Cytoskeleton / metabolism
Female
Meiosis / physiology*
Microtubules / metabolism
Morphogenesis
Nuclear Proteins / metabolism
RNA Interference
Recombinant Fusion Proteins / genetics,  metabolism
Chemical
Reg. No./Substance:
0/Biological Markers; 0/CYB-3 protein, C elegans; 0/Caenorhabditis elegans Proteins; 0/Cell Cycle Proteins; 0/Cullin Proteins; 0/Cyclin B; 0/Cyclin B1; 0/Nuclear Proteins; 0/Recombinant Fusion Proteins; 0/Zyg11 protein, C elegans; 0/cullin-2 protein, C elegans

From MEDLINE®/PubMed®, a database of the U.S. National Library of Medicine


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