Document Detail


Unregulated ARF6 activation in epithelial cysts generates hyperactive signaling endosomes and disrupts morphogenesis.
MedLine Citation:
PMID:  20462959     Owner:  NLM     Status:  MEDLINE    
Abstract/OtherAbstract:
Tumor development in glandular tissues is associated with structural alterations in the hollow ducts and spherical structures that comprise such tissues. We describe a signaling axis involving sustained activation of the GTP-binding protein, ARF6, that provokes dramatic changes in the organization of epithelial cysts, reminiscent of tumorigenic glandular phenotypes. In reconstituted basement membrane cultures of renal epithelial cysts, enhanced ARF6 activation induces the formation of cell-filled glandular structures with multiple lumens and disassembled cadherin-based cell-cell contacts. All of these alterations are accompanied by growth factor receptor internalization into signaling endosomes and reversed by blocking ARF6 activation or receptor endocytosis. Receptor localization in signaling endosomes results in hyperactive extracellular signal-regulated kinase signaling leading to Bcl-2 stabilization and aberrant cysts. Similarly, formation of hyperproliferative and disorganized mammary acini induced by chronic stimulation of colony-stimulating factor 1 receptor is coupled to endogenous ARF6 activation and constitutive receptor internalization and is reversed by ARF6 inhibition. These findings identify a previously unrecognized link between ARF6-regulated receptor internalization and events that drive dramatic alterations in cyst morphogenesis providing new mechanistic insight into the molecular processes that can promote epithelial glandular disruption.
Authors:
Jogender S Tushir; James Clancy; Andrew Warren; Carolyn Wrobel; Joan S Brugge; Crislyn D'Souza-Schorey
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Publication Detail:
Type:  Journal Article; Research Support, N.I.H., Extramural; Research Support, Non-U.S. Gov't     Date:  2010-05-12
Journal Detail:
Title:  Molecular biology of the cell     Volume:  21     ISSN:  1939-4586     ISO Abbreviation:  Mol. Biol. Cell     Publication Date:  2010 Jul 
Date Detail:
Created Date:  2010-06-30     Completed Date:  2010-12-16     Revised Date:  2011-08-01    
Medline Journal Info:
Nlm Unique ID:  9201390     Medline TA:  Mol Biol Cell     Country:  United States    
Other Details:
Languages:  eng     Pagination:  2355-66     Citation Subset:  IM    
Affiliation:
Department of Biological Sciences, University of Notre Dame, Notre Dame, IN 46556, USA.
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MeSH Terms
Descriptor/Qualifier:
ADP-Ribosylation Factors / genetics,  metabolism*
Animals
Cell Line
Dogs
Endosomes / metabolism*
Enzyme Activation
Epithelial Cells / cytology,  metabolism*
Epithelium / anatomy & histology,  embryology*,  metabolism
Extracellular Signal-Regulated MAP Kinases / metabolism
Humans
Mammary Glands, Human / anatomy & histology,  embryology,  metabolism
Morphogenesis / physiology*
Phospholipase D / metabolism
Proto-Oncogene Proteins c-bcl-2 / genetics,  metabolism
Signal Transduction / physiology*
Grant Support
ID/Acronym/Agency:
R01 CA105134-09/CA/NCI NIH HHS
Chemical
Reg. No./Substance:
0/Proto-Oncogene Proteins c-bcl-2; EC 2.7.11.24/Extracellular Signal-Regulated MAP Kinases; EC 3.1.4.4/Phospholipase D; EC 3.6.5.2/ADP-Ribosylation Factors; EC 3.6.5.2/ADP-ribosylation factor 6

From MEDLINE®/PubMed®, a database of the U.S. National Library of Medicine


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