Document Detail


Tight junctions containing claudin 4 and 6 are essential for blastocyst formation in preimplantation mouse embryos.
MedLine Citation:
PMID:  17980358     Owner:  NLM     Status:  MEDLINE    
Abstract/OtherAbstract:
The trophectoderm (TE) is the first epithelium to be generated during mammalian early development. The TE works as a barrier that isolates the inner cell mass from the uterine environment and provides the turgidity of the blastocyst through elevated hydrostatic pressure. In this study, we investigated the role of tight junctions (TJs) in the barrier function of the TE during mouse blastocyst formation. RT-PCR and immunostaining revealed that the mouse TE expressed at least claudin 4, 6, and 7 among the 24 members of the claudin gene family, which encode structural and functional constituents of TJs. When embryos were cultured in the presence of a GST-fused C-terminal half of Clostridium perfringens enterotoxin (GST-C-CPE), a polypeptide with inhibitory activity to claudin 4 and 6, normal blastocyst formation was remarkably inhibited; the embryos had no or an immature blastocoel cavity without expansion, and blastomeres showed a rounded shape. In these embryos, claudin 4 and 6 proteins were absent from TJs and the barrier function of the TE was disrupted; however the basolateral localization of the Na+/K+-ATPase alpha1 subunit and aquaporin 3, which are thought to be involved in blastocyst formation, appeared normal. These results clearly demonstrate that the barrier function of TJs in the TE is required for normal blastocyst formation.
Authors:
Kazumasa Moriwaki; Shoichiro Tsukita; Mikio Furuse
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Publication Detail:
Type:  Journal Article; Research Support, Non-U.S. Gov't     Date:  2007-10-04
Journal Detail:
Title:  Developmental biology     Volume:  312     ISSN:  1095-564X     ISO Abbreviation:  Dev. Biol.     Publication Date:  2007 Dec 
Date Detail:
Created Date:  2007-12-07     Completed Date:  2008-01-23     Revised Date:  -    
Medline Journal Info:
Nlm Unique ID:  0372762     Medline TA:  Dev Biol     Country:  United States    
Other Details:
Languages:  eng     Pagination:  509-22     Citation Subset:  IM    
Affiliation:
Department of Cell Biology, Graduate School of Medicine, Kyoto University, Yoshida-Konoe, Sakyo-ku, Kyoto 606-8501, Japan.
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MeSH Terms
Descriptor/Qualifier:
Animals
Aquaporin 1 / metabolism
Blastocyst / metabolism*
Cell Differentiation
Embryonic Development*
Female
Gene Expression Regulation, Developmental
Membrane Proteins / metabolism*
Mice
Mice, Inbred C57BL
Microscopy, Fluorescence
Recombinant Fusion Proteins / genetics,  metabolism
Sodium-Potassium-Exchanging ATPase / metabolism
Tight Junctions / metabolism*
Chemical
Reg. No./Substance:
0/Membrane Proteins; 0/Recombinant Fusion Proteins; 0/claudin 4; 0/claudin 6; 146410-94-8/Aquaporin 1; EC 3.6.3.9/Sodium-Potassium-Exchanging ATPase

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