Document Detail


Targeted mutagenesis of acyl-lipid desaturases in Synechocystis: evidence for the important roles of polyunsaturated membrane lipids in growth, respiration and photosynthesis.
MedLine Citation:
PMID:  8978669     Owner:  NLM     Status:  MEDLINE    
Abstract/OtherAbstract:
Acyl-lipid desaturases introduce double bonds (unsaturated bonds) at specifically defined positions in fatty acids that are esterified to the glycerol backbone of membrane glycerolipids. The desA, desB and desD genes of Synechocystis sp. PCC 6803 encode acyl-lipid desaturases that introduce double bonds at the delta12, omega3 and delta6 positions of C18 fatty acids respectively. The mutation of each of these genes by insertion of an antibiotic resistance gene cartridge completely eliminated the corresponding desaturation reaction. This system allowed us to manipulate the number of unsaturated bonds in membrane glycerolipids in this organism in a step-wise manner. Comparisons of the variously mutated cells revealed that the replacement of all polyunsaturated fatty acids by a monounsaturated fatty acid suppressed growth of the cells at low temperature and, moreover, it decreased the tolerance of the cells to photoinhibition of photosynthesis at low temperature by suppressing recovery of the photosystem II protein complex from photoinhibitory damage. However, the replacement of tri- and tetraunsaturated fatty acids by a diunsaturated fatty acid did not have such effects. These findings indicate that polyunsaturated fatty acids are important in protecting the photosynthetic machinery from photoinhibition at low temperatures.
Authors:
Y Tasaka; Z Gombos; Y Nishiyama; P Mohanty; T Ohba; K Ohki; N Murata
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Publication Detail:
Type:  Journal Article; Research Support, Non-U.S. Gov't    
Journal Detail:
Title:  The EMBO journal     Volume:  15     ISSN:  0261-4189     ISO Abbreviation:  EMBO J.     Publication Date:  1996 Dec 
Date Detail:
Created Date:  1997-01-27     Completed Date:  1997-01-27     Revised Date:  2010-09-13    
Medline Journal Info:
Nlm Unique ID:  8208664     Medline TA:  EMBO J     Country:  ENGLAND    
Other Details:
Languages:  eng     Pagination:  6416-25     Citation Subset:  IM    
Affiliation:
Department of Regulation Biology, National Institute for Basic Biology, Myodaiji, Okazaki, Japan.
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MeSH Terms
Descriptor/Qualifier:
Calorimetry, Differential Scanning
Chlorophyll / metabolism
Cyanobacteria / enzymology*,  genetics,  growth & development
Fatty Acid Desaturases / genetics,  metabolism*
Fatty Acids, Unsaturated / metabolism*
Genes, Bacterial
Glycerides / metabolism
Kinetics
Membrane Lipids / metabolism*
Mutagenesis, Insertional
Oxygen Consumption*
Photosynthesis*
Plasmids
Recombinant Proteins / metabolism
Temperature
Chemical
Reg. No./Substance:
0/Fatty Acids, Unsaturated; 0/Glycerides; 0/Membrane Lipids; 0/Recombinant Proteins; 1406-65-1/Chlorophyll; EC 1.14.19.-/Fatty Acid Desaturases; EC 1.14.99.-/delta-9 fatty acid desaturase
Comments/Corrections

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