Document Detail


TGF-beta regulates T-cell neurokinin-1 receptor internalization and function.
MedLine Citation:
PMID:  20160079     Owner:  NLM     Status:  MEDLINE    
Abstract/OtherAbstract:
Substance P (SP) is a proinflammatory mediator implicated in inflammatory bowel disease (IBD) and other inflammatory states. SP acts by stimulating the neurokinin-1 receptor (NK-1R) on T lymphocytes and other cell types, and regulates these cells in a complex interplay with multiple cytokines. The mechanisms of interaction among these inflammatory mediators are not yet fully understood. Here, we demonstrate that function of the NK-1R, a member of the G protein-coupled receptor (GPCR) superfamily, is modulated by TGF-beta. The latter acts not on a GPCR but via serine-threonine kinase-class receptors. By flow confocal image analysis, we demonstrate that TGF-beta delays SP-induced NK-1R internalization on mucosal T cells isolated from a mouse model of IBD and on granuloma T cells in murine schistosomiasis. Furthermore, luciferase reporter-gene assays revealed that NK-1R stimulation activates the nuclear factor of activated T cell- and activator protein-1-dependent signaling pathways, which are known triggers of effector T-cell cytokine production. TGF-beta markedly increases SP-induced activation of these signaling cascades, suggesting that delayed NK-1R internalization results in enhanced signaling. Providing a link to amplified immune function, SP and TGF-beta, when applied in combination, trigger a strong release of the proinflammatory cytokines IFN-gamma and IL17 from intestinal inflammatory T cells, whereas either agonist alone shows no effect. These observations establish precedent that members of two distinct receptor superfamilies can interact via a previously unrecognized mechanism, and reveal a paradigm of GPCR transregulation that is relevant to IBD and possibly other disease processes.
Authors:
Martin Beinborn; Arthur Blum; Long Hang; Tommy Setiawan; Jonathan C Schroeder; Korynn Stoyanoff; John Leung; Joel V Weinstock
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Publication Detail:
Type:  Journal Article; Research Support, N.I.H., Extramural; Research Support, Non-U.S. Gov't     Date:  2010-02-16
Journal Detail:
Title:  Proceedings of the National Academy of Sciences of the United States of America     Volume:  107     ISSN:  1091-6490     ISO Abbreviation:  Proc. Natl. Acad. Sci. U.S.A.     Publication Date:  2010 Mar 
Date Detail:
Created Date:  2010-03-03     Completed Date:  2010-04-16     Revised Date:  2010-09-03    
Medline Journal Info:
Nlm Unique ID:  7505876     Medline TA:  Proc Natl Acad Sci U S A     Country:  United States    
Other Details:
Languages:  eng     Pagination:  4293-8     Citation Subset:  IM    
Affiliation:
Division of Gastroenterology and Hepatology, Department of Internal Medicine, and Molecular Pharmacology Research Center, Molecular Cardiology Research Institute, Tufts Medical Center, Boston, MA 02111, USA.
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MeSH Terms
Descriptor/Qualifier:
Animals
Cell Line
Endocytosis*
Flow Cytometry
Humans
Inflammatory Bowel Diseases / immunology
Interleukin-10 / genetics,  physiology
Mice
Mice, Inbred C57BL
Microscopy, Confocal
Receptors, Neurokinin-1 / immunology*,  metabolism
Signal Transduction
Substance P
T-Lymphocytes / immunology*
Transforming Growth Factor beta / physiology*
Grant Support
ID/Acronym/Agency:
DK058755/DK/NIDDK NIH HHS; DK38327/DK/NIDDK NIH HHS
Chemical
Reg. No./Substance:
0/Receptors, Neurokinin-1; 0/Transforming Growth Factor beta; 130068-27-8/Interleukin-10; 33507-63-0/Substance P

From MEDLINE®/PubMed®, a database of the U.S. National Library of Medicine


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