Document Detail


Syntaxin 1A promotes the endocytic sorting of EAAC1 leading to inhibition of glutamate transport.
MedLine Citation:
PMID:  16959903     Owner:  NLM     Status:  MEDLINE    
Abstract/OtherAbstract:
The neuronal glutamate transporter, excitatory amino-acid carrier 1 (EAAC1), plays an important role in the modulation of neurotransmission and contributes to synthesis of the inhibitory neurotransmitter gamma-aminobutyric acid (GABA) and to epileptogenesis. However, the mechanisms that regulate EAAC1 endocytic sorting and function remain largely unknown. Here, we first demonstrate that EAAC1 undergoes internalization through the clathrin-mediated pathway and further show that syntaxin 1A, a key molecule in synaptic exocytosis, potentiates EAAC1 internalization, thus leading to the functional inhibition of EAAC1. In the presence of the transmembrane domain of syntaxin 1A, its H3 coiled-coil domain of syntaxin 1A is necessary and sufficient for the inhibition of EAAC1. Furthermore, specific suppression of endogenous syntaxin 1A significantly blocked EAAC1 endocytic sorting and lysosomal degradation promoted by kainic acid, a drug for kindling the animal model of human temporal lobe epilepsy in rat, indicating a potential role of syntaxin 1A in epileptogenesis. These findings provide new evidence that syntaxin 1A serves as an intrinsic enhancer to EAAC1 endocytic sorting and further suggest that syntaxin 1A is conversant with both ;ins' and ;outs' of synaptic neurotransmission.
Authors:
Yong-Xin Yu; Li Shen; Peng Xia; Ya-Wei Tang; Lan Bao; Gang Pei
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Publication Detail:
Type:  Journal Article; Research Support, Non-U.S. Gov't    
Journal Detail:
Title:  Journal of cell science     Volume:  119     ISSN:  0021-9533     ISO Abbreviation:  J. Cell. Sci.     Publication Date:  2006 Sep 
Date Detail:
Created Date:  2006-09-08     Completed Date:  2006-12-08     Revised Date:  -    
Medline Journal Info:
Nlm Unique ID:  0052457     Medline TA:  J Cell Sci     Country:  England    
Other Details:
Languages:  eng     Pagination:  3776-87     Citation Subset:  IM    
Affiliation:
Laboratory of Molecular Cell Biology, Institute of Biochemistry and Cell Biology, Shanghai Institutes of Biological Sciences, Chinese Academy of Sciences, Shanghai, People's Republic of China.
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MeSH Terms
Descriptor/Qualifier:
Amino Acid Transport Systems / drug effects,  metabolism*
Animals
Biological Transport
Cells, Cultured
Clathrin / metabolism
Coated Pits, Cell-Membrane / drug effects,  metabolism
Endocytosis / drug effects,  physiology*
Endosomes / drug effects
Excitatory Amino Acid Transporter 3 / genetics,  metabolism*
Gene Expression Regulation / drug effects
Glutamates / metabolism*
Humans
Kainic Acid / pharmacology
Lysosomes / drug effects
Neurons / drug effects
Protein Binding / drug effects
Protein Structure, Tertiary
RNA, Messenger / genetics,  metabolism
Rats
Rats, Sprague-Dawley
Syntaxin 1 / chemistry,  metabolism*
Chemical
Reg. No./Substance:
0/Amino Acid Transport Systems; 0/Clathrin; 0/Excitatory Amino Acid Transporter 3; 0/Glutamates; 0/RNA, Messenger; 0/Slc1a1 protein, rat; 0/Stx1a protein, rat; 0/Syntaxin 1; 487-79-6/Kainic Acid

From MEDLINE®/PubMed®, a database of the U.S. National Library of Medicine


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