Document Detail


Substrate filtering by the active site crossover loop in UCHL3 revealed by sortagging and gain-of-function mutations.
MedLine Citation:
PMID:  19047059     Owner:  NLM     Status:  MEDLINE    
Abstract/OtherAbstract:
Determining how deubiquitinating enzymes discriminate between ubiquitin-conjugated substrates is critical to understand their function. Through application of a novel protein cleavage and tagging technique, sortagging, we show that human UCHL3 and the Plasmodium falciparum homologue, members of the ubiquitin C-terminal hydrolase family, use a unique active site crossover loop to restrict access of bulky ubiquitin adducts to the active site. Although it provides connectivity for critical active site residues in UCHL3, physical integrity of the crossover loop is dispensable for catalysis. By enlarging the active site crossover loop, we have constructed gain-of-function mutants that can accept substrates that the parent enzyme cannot, including ubiquitin chains of various linkages.
Authors:
Maximilian W Popp; Katerina Artavanis-Tsakonas; Hidde L Ploegh
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Publication Detail:
Type:  Journal Article; Research Support, N.I.H., Extramural     Date:  2008-12-01
Journal Detail:
Title:  The Journal of biological chemistry     Volume:  284     ISSN:  0021-9258     ISO Abbreviation:  J. Biol. Chem.     Publication Date:  2009 Feb 
Date Detail:
Created Date:  2009-02-02     Completed Date:  2009-04-06     Revised Date:  2010-09-23    
Medline Journal Info:
Nlm Unique ID:  2985121R     Medline TA:  J Biol Chem     Country:  United States    
Other Details:
Languages:  eng     Pagination:  3593-602     Citation Subset:  IM    
Affiliation:
Whitehead Institute for Biomedical Research, Cambridge, Massachusetts 02142, USA.
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MeSH Terms
Descriptor/Qualifier:
Animals
Catalysis
Catalytic Domain / physiology
Cysteine Endopeptidases / chemistry*,  genetics,  metabolism
Humans
Mutation
Plasmodium falciparum / enzymology*,  genetics,  metabolism
Protein Structure, Secondary / physiology
Protozoan Proteins / chemistry*,  metabolism
Substrate Specificity / physiology
Ubiquitin / chemistry*,  genetics,  metabolism
Ubiquitination / physiology*
Chemical
Reg. No./Substance:
0/Protozoan Proteins; 0/Ubiquitin; EC 3.1.2.15/UCHL3 protein, human; EC 3.4.22.-/Cysteine Endopeptidases
Comments/Corrections

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