Document Detail


Second messenger-mediated spatiotemporal control of protein degradation regulates bacterial cell cycle progression.
MedLine Citation:
PMID:  19136627     Owner:  NLM     Status:  MEDLINE    
Abstract/OtherAbstract:
Second messengers control a wide range of important cellular functions in eukaryotes and prokaryotes. Here we show that cyclic di-GMP, a global bacterial second messenger, promotes cell cycle progression in Caulobacter crescentus by mediating the specific degradation of the replication initiation inhibitor CtrA. During the G1-to-S-phase transition, both CtrA and its cognate protease ClpXP dynamically localize to the old cell pole, where CtrA is rapidly degraded. Sequestration of CtrA to the cell pole depends on PopA, a newly identified cyclic di-GMP effector protein. PopA itself localizes to the cell pole and directs CtrA to this subcellular site via the direct interaction with a mediator protein, RcdA. We present evidence that c-di-GMP regulates CtrA degradation during the cell cycle by controlling the dynamic sequestration of the PopA recruitment factor to the cell pole. Furthermore, we show that cell cycle timing of CtrA degradation relies on converging pathways responsible for substrate and protease localization to the old cell pole. This is the first report that links cyclic di-GMP to protein dynamics and cell cycle control in bacteria.
Authors:
Anna Duerig; Sören Abel; Marc Folcher; Micael Nicollier; Torsten Schwede; Nicolas Amiot; Bernd Giese; Urs Jenal
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Publication Detail:
Type:  Journal Article; Research Support, Non-U.S. Gov't    
Journal Detail:
Title:  Genes & development     Volume:  23     ISSN:  1549-5477     ISO Abbreviation:  Genes Dev.     Publication Date:  2009 Jan 
Date Detail:
Created Date:  2009-01-12     Completed Date:  2009-01-30     Revised Date:  2009-11-18    
Medline Journal Info:
Nlm Unique ID:  8711660     Medline TA:  Genes Dev     Country:  United States    
Other Details:
Languages:  eng     Pagination:  93-104     Citation Subset:  IM    
Affiliation:
Biozentrum, University of Basel, 4056 Basel, Switzerland.
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MeSH Terms
Descriptor/Qualifier:
Bacterial Proteins / metabolism*
Caulobacter crescentus / genetics*,  metabolism*
Cell Cycle / physiology*
DNA-Binding Proteins / metabolism
Protein Binding
Protein Transport / physiology
Second Messenger Systems / physiology*
Time Factors
Transcription Factors / metabolism
Chemical
Reg. No./Substance:
0/Bacterial Proteins; 0/CtrA protein, Caulobacter; 0/DNA-Binding Proteins; 0/PleD protein, Caulobacter crescentus; 0/Transcription Factors
Comments/Corrections

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