Document Detail


Sam68 and ERKs regulate leptin-induced expression of OB-Rb mRNA in C2C12 myotubes.
MedLine Citation:
PMID:  19524014     Owner:  NLM     Status:  MEDLINE    
Abstract/OtherAbstract:
Acute leptin treatment significantly increases the mRNA of the long isoform of leptin receptor (OB-Rb) in C2C12 myotubes after as little as 30min, without affecting that of the short isoform (OB-Ra). The Sam68 STAR protein has been implicated in leptin signal transduction as an adaptor molecule useful to recruit other signalling proteins. We found that leptin increased Sam68 tyrosine-phosphorylation so decreasing its poly(U)-binding capacity. RT-PCR analysis of the mRNA bound to immunoprecipitated Sam68 showed that Sam68 associated with OB-Rb but not OB-Ra mRNA in control and leptin-treated C2C12 cells. The siRNA-mediated silencing of Sam68 reduced its levels by 89% and abolished the leptin-mediated increase in OB-Rb mRNA. Leptin activates ERKs which in turn might phosphorylate Sam68 modifying its influence on mRNA. We did not observe any changes in Sam68 Ser/Thr phosphorylation but using the specific MEK1 inhibitor PD-98059 showed that leptin-mediated ERK activation is essential for leptin's effect on OB-Rb mRNA expression. Thus it appears that leptin has a positive short-term effect on the regulation of OB-Rb mRNA in C2C12 cells, involving both Sam68 and ERKs. These results might suggest that leptin signal acutely favours its own sensitivity.
Authors:
P Maroni; L Citterio; R Piccoletti; P Bendinelli
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Publication Detail:
Type:  Journal Article; Research Support, Non-U.S. Gov't     Date:  2009-06-11
Journal Detail:
Title:  Molecular and cellular endocrinology     Volume:  309     ISSN:  1872-8057     ISO Abbreviation:  Mol. Cell. Endocrinol.     Publication Date:  2009 Oct 
Date Detail:
Created Date:  2009-08-03     Completed Date:  2009-10-26     Revised Date:  2009-11-19    
Medline Journal Info:
Nlm Unique ID:  7500844     Medline TA:  Mol Cell Endocrinol     Country:  Ireland    
Other Details:
Languages:  eng     Pagination:  26-31     Citation Subset:  IM    
Affiliation:
Istituto Ortopedico Galeazzi, IRCCS, Milan, Italy.
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MeSH Terms
Descriptor/Qualifier:
Adaptor Proteins, Signal Transducing / metabolism*
Animals
Extracellular Signal-Regulated MAP Kinases / antagonists & inhibitors,  metabolism*
Gene Expression Regulation / drug effects*
Humans
Leptin / pharmacology*
Mice
Muscle Fibers, Skeletal / drug effects*,  enzymology*
Phosphotyrosine / metabolism
Poly U / metabolism
Protein Isoforms / genetics,  metabolism
Protein Kinase Inhibitors / pharmacology
RNA, Messenger / genetics,  metabolism
RNA-Binding Proteins / metabolism*
Receptors, Leptin / genetics*,  metabolism
Chemical
Reg. No./Substance:
0/Adaptor Proteins, Signal Transducing; 0/Khdrbs1 protein, mouse; 0/Leptin; 0/Protein Isoforms; 0/Protein Kinase Inhibitors; 0/RNA, Messenger; 0/RNA-Binding Proteins; 0/Receptors, Leptin; 21820-51-9/Phosphotyrosine; 27416-86-0/Poly U; EC 2.7.11.24/Extracellular Signal-Regulated MAP Kinases

From MEDLINE®/PubMed®, a database of the U.S. National Library of Medicine


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