Document Detail


Role of the von Hippel-Lindau tumour suppressor protein in the regulation of HIF-1alpha and its oxygen-regulated transactivation domains at high cell density.
MedLine Citation:
PMID:  15750626     Owner:  NLM     Status:  MEDLINE    
Abstract/OtherAbstract:
Hypoxia-inducible factor-1alpha (HIF-1alpha) induction and associated transcription were investigated during high cell density, focusing on the negative regulator of HIF-1alpha expression, the von Hippel-Lindau (VHL) protein. In 293T and HeLa cells, HIF-1alpha protein levels and associated transcription were induced as cells approached confluence. To determine whether these changes were due to a deficit in nuclear VHL-mediated ubiquitination of HIF-1alpha at confluence, cells were stably transfected with VHL. Overexpression of VHL in 293T cells had no demonstrable effect on the induction and nuclear accumulation of HIF-1alpha during high cell density or associated transcription. Moreover, RCC cells stably transfected with full-length VHL failed to exhibit the cell-density-dependent induction of HIF-1alpha noted in other cell lines. Investigation of both N-terminal and C-terminal (aa 727-826) oxygen-regulated proline and asparagine hydroxylation of HIF-1alpha revealed that both are inhibited during high cell density, as determined by impaired capture of HIF-1alpha by VHL and enhanced C-terminal transactivation. Finally, cell-density-mediated induction of HIF-1alpha and GLUT1 in RCC cells could be completely reconstituted by mutations in VHL binding affinity, suggesting that cell-density dependent induction of HIF-1alpha and transactivation may underpin some of the deregulated gene expression observed in VHL disease.
Authors:
Steve M Paltoglou; Ben J Roberts
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Publication Detail:
Type:  Journal Article; Research Support, Non-U.S. Gov't    
Journal Detail:
Title:  Oncogene     Volume:  24     ISSN:  0950-9232     ISO Abbreviation:  Oncogene     Publication Date:  2005 May 
Date Detail:
Created Date:  2005-05-26     Completed Date:  2005-06-09     Revised Date:  2008-11-21    
Medline Journal Info:
Nlm Unique ID:  8711562     Medline TA:  Oncogene     Country:  England    
Other Details:
Languages:  eng     Pagination:  3830-5     Citation Subset:  IM    
Affiliation:
School of Pharmaceutical, Molecular and Biomedical Sciences, University of South Australia, Reid Building, Frome Rd., Adelaide 5000, Australia.
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MeSH Terms
Descriptor/Qualifier:
Cell Count
Cell Line
Glucose Transporter Type 1
Humans
Hypoxia-Inducible Factor 1, alpha Subunit
Monosaccharide Transport Proteins / biosynthesis
Response Elements / physiology
Transcription Factors / biosynthesis*
Transcriptional Activation*
Tumor Suppressor Proteins / physiology*
Ubiquitin-Protein Ligases / physiology*
Von Hippel-Lindau Tumor Suppressor Protein
Chemical
Reg. No./Substance:
0/Glucose Transporter Type 1; 0/HIF1A protein, human; 0/Hypoxia-Inducible Factor 1, alpha Subunit; 0/Monosaccharide Transport Proteins; 0/SLC2A1 protein, human; 0/Transcription Factors; 0/Tumor Suppressor Proteins; EC 6.3.2.19/Ubiquitin-Protein Ligases; EC 6.3.2.19/VHL protein, human; EC 6.3.2.19/Von Hippel-Lindau Tumor Suppressor Protein

From MEDLINE®/PubMed®, a database of the U.S. National Library of Medicine


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