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Rod, M-cone and M/S-cone inputs to hyperpolarizing bipolar cells in the mouse retina.
MedLine Citation:
PMID:  22219344     Owner:  NLM     Status:  Publisher    
Abstract/OtherAbstract:
Bipolar cells are the central neurons of the retina that convey visual signals from rod and cone photoreceptors in the outer retina to higher-order neurons in the inner retina and the brain. Early anatomical studies have suggested that there are 4 types of cone hyperpolarizing (OFF) bipolar cells (HBCs) in the mouse retina, but no light responses have been systematically examined. By analyzing light-evoked cation and chloride currents (ΔIC and ΔICl) from over 50 morphologically-identified HBCs in the dark-adapted wildtype and connexin36 knockout (Cx36-/-) mouse retinas, we identified three types of HBCs, each with distinct light responses and morphological characteristics. The HBCR/MCs with axon terminals ramifying between 0% and 30% of the inner plexiform layer (IPL) receive mixed inputs from rods and M-cones, the HBCMCs with axon terminals ramifying between 10% and 50% of the IPL receive inputs primarily from M-cones, and the HBCM/SCs with axon terminals ramifying between 25% and 50% of IPL receive inputs primarily from cones with mixed M- and S-cone pigments. Moreover, we found that HBCR/MCs in the Cx36-/- mice exhibit light responses very similar to the wildtype HBCR/MCs, suggesting that the mixed rod/cone inputs are not mediated by connexin36-dependent rod-cone coupling, but rather by direct synaptic contacts from rods and M-cones. This study constitutes the first systematic investigation that correlates light response characteristics and axonal morphology of HBCs in dark-adapted mouse retina, and contributes to recently emerging evidence that revises the traditional view that mammalian HBCs only contact cone photoreceptors.
Authors:
Ji-Jie Pang; Fan Gao; David L Paul; Samuel M Wu
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Publication Detail:
Type:  JOURNAL ARTICLE     Date:  2012-1-4
Journal Detail:
Title:  The Journal of physiology     Volume:  -     ISSN:  1469-7793     ISO Abbreviation:  -     Publication Date:  2012 Jan 
Date Detail:
Created Date:  2012-1-5     Completed Date:  -     Revised Date:  -    
Medline Journal Info:
Nlm Unique ID:  0266262     Medline TA:  J Physiol     Country:  -    
Other Details:
Languages:  ENG     Pagination:  -     Citation Subset:  -    
Affiliation:
Baylor College of Medicine;
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