Document Detail


Regulation of the subcellular localization of cyclic AMP-dependent protein kinase in response to physiological stresses and sexual differentiation in the fission yeast Schizosaccharomyces pombe.
MedLine Citation:
PMID:  18621924     Owner:  NLM     Status:  MEDLINE    
Abstract/OtherAbstract:
We describe regulation of the subcellular localization of cyclic AMP (cAMP)-dependent protein kinase (PKA) regulatory (Cgs1p) and catalytic (Pka1p) subunits in the fission yeast Schizosaccharomyces pombe in response to physiological stresses and during sexual differentiation as determined by fluorescence microscopy of the Cgs1-green fluorescent protein (GFP) and Pka1-GFP fusion proteins, respectively. In wild-type S. pombe cells cultured to log phase under normal growth conditions, Cgs1p and Pka1p are concentrated in the nucleus and more diffusely present in the cytoplasm. Nuclear localization of both proteins is dependent on cAMP, since in cells lacking adenylate cyclase they are detectable only in the cytoplasm. In cells lacking Cgs1p or both Cgs1p and adenylate cyclase, Pka1p is concentrated in the nucleus, demonstrating a role for Cgs1p in the nuclear exclusion of Pka1p. Nuclear-cytoplasmic redistribution of Cgs1p and Pka1p is triggered by growth in glucose-limited or hyperosmotic media and in response to stationary-phase growth. In addition, both proteins are excluded from the nucleus in mating cells undergoing karyogamy and subsequently concentrated in postmeiotic spores. Cgs1p is required for subcellular redistribution of Pka1p induced by growth in glucose-limited and hyperosmotic media and during karyogamy but is not required for Pka1p redistribution triggered by stationary-phase growth or for the enrichment of Pka1p in spores. Our results demonstrate that PKA localization is regulated by cAMP and regulatory subunit-dependent and -independent mechanisms in S. pombe.
Authors:
Yasuhiro Matsuo; Brittney McInnis; Stevan Marcus
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Publication Detail:
Type:  Journal Article; Research Support, N.I.H., Extramural     Date:  2008-07-11
Journal Detail:
Title:  Eukaryotic cell     Volume:  7     ISSN:  1535-9786     ISO Abbreviation:  Eukaryotic Cell     Publication Date:  2008 Sep 
Date Detail:
Created Date:  2008-09-08     Completed Date:  2008-10-30     Revised Date:  2009-11-18    
Medline Journal Info:
Nlm Unique ID:  101130731     Medline TA:  Eukaryot Cell     Country:  United States    
Other Details:
Languages:  eng     Pagination:  1450-9     Citation Subset:  IM    
Affiliation:
Department of Biological Sciences, The University of Alabama, 343A Shelby Hall, 250 Hackberry Lane, Tuscaloosa, AL 35487, USA.
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MeSH Terms
Descriptor/Qualifier:
Cell Nucleus / enzymology,  genetics,  physiology
Cyclic AMP / metabolism
Cyclic AMP-Dependent Protein Kinases / genetics,  metabolism*
Cytoplasm / enzymology,  genetics,  physiology
Gene Expression Regulation, Enzymologic*
Gene Expression Regulation, Fungal*
Glucose / metabolism
Osmolar Concentration
Protein Subunits / genetics,  metabolism*
Protein Transport
Schizosaccharomyces / enzymology*,  genetics,  physiology*
Schizosaccharomyces pombe Proteins / genetics,  metabolism*
Grant Support
ID/Acronym/Agency:
R01GM068685/GM/NIGMS NIH HHS
Chemical
Reg. No./Substance:
0/Protein Subunits; 0/Schizosaccharomyces pombe Proteins; 50-99-7/Glucose; 60-92-4/Cyclic AMP; EC 2.7.11.11/Cyclic AMP-Dependent Protein Kinases
Comments/Corrections

From MEDLINE®/PubMed®, a database of the U.S. National Library of Medicine


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