Document Detail


Recombination proteins mediate meiotic spatial chromosome organization and pairing.
MedLine Citation:
PMID:  20371348     Owner:  NLM     Status:  MEDLINE    
Abstract/OtherAbstract:
Meiotic chromosome pairing involves not only recognition of homology but also juxtaposition of entire chromosomes in a topologically regular way. Analysis of filamentous fungus Sordaria macrospora reveals that recombination proteins Mer3, Msh4, and Mlh1 play direct roles in all of these aspects, in advance of their known roles in recombination. Absence of Mer3 helicase results in interwoven chromosomes, thereby revealing the existence of features that specifically ensure "entanglement avoidance." Entanglements that remain at zygotene, i.e., "interlockings," require Mlh1 for resolution, likely to eliminate constraining recombinational connections. Patterns of Mer3 and Msh4 foci along aligned chromosomes show that the double-strand breaks mediating homologous alignment have spatially separated ends, one localized to each partner axis, and that pairing involves interference among developing interhomolog interactions. We propose that Mer3, Msh4, and Mlh1 execute all of these roles during pairing by modulating the state of nascent double-strand break/partner DNA contacts within axis-associated recombination complexes.
Authors:
Aurora Storlazzi; Silvana Gargano; Gwenael Ruprich-Robert; Matthieu Falque; Michelle David; Nancy Kleckner; Denise Zickler
Related Documents :
19099188 - Chromatin structure contribution to the synaptonemal complex formation.
8243168 - Two new x-autosome robertsonian translocations in the mouse. ii. sex chromosome configu...
10101178 - Distribution of crossing over on mouse synaptonemal complexes using immunofluorescent l...
10973068 - The pachytene checkpoint.
18266248 - Array comparative genomic hybridization (acgh) analysis in prader-willi syndrome.
17253988 - The fluorouridine insensitive 1 (fur1) mutant is defective in equilibrative nucleoside ...
Publication Detail:
Type:  Journal Article; Research Support, N.I.H., Extramural; Research Support, Non-U.S. Gov't    
Journal Detail:
Title:  Cell     Volume:  141     ISSN:  1097-4172     ISO Abbreviation:  Cell     Publication Date:  2010 Apr 
Date Detail:
Created Date:  2010-04-07     Completed Date:  2010-04-22     Revised Date:  2011-07-27    
Medline Journal Info:
Nlm Unique ID:  0413066     Medline TA:  Cell     Country:  United States    
Other Details:
Languages:  eng     Pagination:  94-106     Citation Subset:  IM    
Copyright Information:
Copyright 2010 Elsevier Inc. All rights reserved.
Affiliation:
Institut de Génétique et Microbiologie, UMR 8621, Université Paris-Sud, 91405 Orsay, France.
Export Citation:
APA/MLA Format     Download EndNote     Download BibTex
MeSH Terms
Descriptor/Qualifier:
Cell Cycle Proteins / metabolism*
Chromosome Pairing*
Fungal Proteins / metabolism*
Meiosis*
Sordariales / cytology*,  metabolism*
Grant Support
ID/Acronym/Agency:
GM025326/GM/NIGMS NIH HHS; GM044794/GM/NIGMS NIH HHS; R01 GM025326-28/GM/NIGMS NIH HHS; R01 GM025326-29/GM/NIGMS NIH HHS; R01 GM025326-33/GM/NIGMS NIH HHS; R01 GM044794-19/GM/NIGMS NIH HHS; R01 GM044794-20S1/GM/NIGMS NIH HHS
Chemical
Reg. No./Substance:
0/Cell Cycle Proteins; 0/Fungal Proteins
Comments/Corrections

From MEDLINE®/PubMed®, a database of the U.S. National Library of Medicine


Previous Document:  Persistent telomere damage induces bypass of mitosis and tetraploidy.
Next Document:  Dynamic and static interactions between p120 catenin and E-cadherin regulate the stability of cell-c...