Document Detail


Rat gonadotropin-releasing hormone receptor expressed in insect cells induces activation of adenylyl cyclase.
MedLine Citation:
PMID:  9484907     Owner:  NLM     Status:  MEDLINE    
Abstract/OtherAbstract:
Increasing evidence exist that multiple G proteins mediate the effects of gonadotropin-releasing hormone (GnRH) on the synthesis and release of pituitary gonadotropins. In the present study, we have expressed the rat GnRH receptor (GnRH-R) in insect cells, by infection with a recombinant baculovirus. Under the conditions used, insect cells expressed, 48 h post-infection, a maximum of 7800 +/- 650 receptors/cell which bound GnRH agonist [D-Trp6]GnRH with a Kd = 0.52 +/- 0.06 nM indicating characteristics similar to those of the natural receptor. No binding was observed in non-infected cells or cells infected with wild-type baculovirus. In presence of GnRH, GnRH-R expressing cells elicited a time- and dose-dependent production of inositol trisphosphate, with a maximum level reached within 30 min and an EC50 = 5 nM. These recombinant insect cells also produced cAMP in response to GnRH. However, in contrast to other heterologous systems, or rat pituitary gonadotropes wherein GnRH induced a weak and delayed elevation of cAMP, in insect cells the rise of cAMP was comparatively rapid, attaining a maximum level after 2 h, and the EC50 was 5 nM. Finally, a clear activation of adenylyl cyclase (AC) in response to GnRH was shown for the first time by measuring the conversion of [alpha-32P]ATP into labeled cAMP, using membrane preparations from GnRH-R expressing insect cells. These data demonstrate that rat GnRH-R has the potential for dual coupling to both phosphoinositidase C and AC and suggest a major influence of the host cell for this coupling and/or its expression, probably in relation with the G protein repertoire and preference. This notion could be extended to several target cells other than pituitary gonadotropes that normally express the GnRH-R in mammals, including hippocampal, Leydig, granulosa, placental and GnRH-secreting hypothalamic cells.
Authors:
R Delahaye; P R Manna; A Bérault; J Berreur-Bonnenfant; P Berreur; R Counis
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Publication Detail:
Type:  Journal Article; Research Support, Non-U.S. Gov't    
Journal Detail:
Title:  Molecular and cellular endocrinology     Volume:  135     ISSN:  0303-7207     ISO Abbreviation:  Mol. Cell. Endocrinol.     Publication Date:  1997 Dec 
Date Detail:
Created Date:  1998-04-08     Completed Date:  1998-04-08     Revised Date:  2006-11-15    
Medline Journal Info:
Nlm Unique ID:  7500844     Medline TA:  Mol Cell Endocrinol     Country:  IRELAND    
Other Details:
Languages:  eng     Pagination:  119-27     Citation Subset:  IM    
Affiliation:
Endocrinologie Cellulaire et Moléculaire de la Reproduction, Université P. & M. Curie, URA CNRS 1449, Paris, France.
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MeSH Terms
Descriptor/Qualifier:
Adenylate Cyclase / metabolism*
Animals
Cell Line
Clone Cells / cytology,  metabolism
Cyclic AMP / metabolism
Enzyme Activation / drug effects,  physiology
Gene Expression / genetics,  physiology
Inositol 1,4,5-Trisphosphate / metabolism
Insects / cytology,  genetics,  metabolism
Protein Binding
Rats
Receptors, LHRH / genetics*,  metabolism,  physiology
Recombinant Proteins / genetics,  metabolism,  pharmacology
Chemical
Reg. No./Substance:
0/Receptors, LHRH; 0/Recombinant Proteins; 60-92-4/Cyclic AMP; 85166-31-0/Inositol 1,4,5-Trisphosphate; EC 4.6.1.1/Adenylate Cyclase

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