Document Detail


Rab conversion as a mechanism of progression from early to late endosomes.
MedLine Citation:
PMID:  16143105     Owner:  NLM     Status:  MEDLINE    
Abstract/OtherAbstract:
The mechanisms of endosome biogenesis and maintenance are largely unknown. The small GTPases Rab 5 and Rab 7 are key determinants of early and late endosomes, organizing effector proteins into specific membrane subdomains. Whether such Rab machineries are indefinitely maintained on membranes or can disassemble in the course of cargo transport is an open question. Here, we combined novel image-analysis algorithms with fast live-cell imaging. We found that the level of Rab 5 dynamically fluctuates on individual early endosomes, linked by fusion and fission events into a network in time. Within it, degradative cargo concentrates in progressively fewer and larger endosomes that migrate from the cell periphery to the center where Rab 5 is rapidly replaced with Rab 7. The class C VPS/HOPS complex, an established GEF for Rab 7, interacts with Rab 5 and is required for Rab 5-to-Rab 7 conversion. Our results reveal unexpected dynamics of Rab domains and suggest Rab conversion as the mechanism of cargo progression between early and late endosomes.
Authors:
Jochen Rink; Eric Ghigo; Yannis Kalaidzidis; Marino Zerial
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Publication Detail:
Type:  Journal Article; Research Support, Non-U.S. Gov't    
Journal Detail:
Title:  Cell     Volume:  122     ISSN:  0092-8674     ISO Abbreviation:  Cell     Publication Date:  2005 Sep 
Date Detail:
Created Date:  2005-09-06     Completed Date:  2005-11-08     Revised Date:  2006-11-15    
Medline Journal Info:
Nlm Unique ID:  0413066     Medline TA:  Cell     Country:  United States    
Other Details:
Languages:  eng     Pagination:  735-49     Citation Subset:  IM    
Affiliation:
Max Planck Institute of Molecular Cell Biology and Genetics, Pfotenhauerstrasse 108, 01307 Dresden, Germany.
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MeSH Terms
Descriptor/Qualifier:
Biological Transport
Cell Line, Tumor
Endosomes / metabolism*
Fluorescent Dyes
Humans
Image Processing, Computer-Assisted
Lipoproteins, LDL / metabolism
Microscopy, Video
Models, Biological
rab GTP-Binding Proteins / metabolism*
rab5 GTP-Binding Proteins / metabolism
Chemical
Reg. No./Substance:
0/Fluorescent Dyes; 0/Lipoproteins, LDL; 152989-05-4/rab7 protein; EC 3.6.1.-/rab GTP-Binding Proteins; EC 3.6.5.2/rab5 GTP-Binding Proteins

From MEDLINE®/PubMed®, a database of the U.S. National Library of Medicine


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