Document Detail


RNA-binding protein HuR regulates RGS4 mRNA stability in rabbit colonic smooth muscle cells.
MedLine Citation:
PMID:  20881234     Owner:  NLM     Status:  MEDLINE    
Abstract/OtherAbstract:
Regulator of G protein signaling 4 (RGS4) regulates the strength and duration of G protein signaling and plays an important role in smooth muscle contraction, cardiac development, and psychiatric disorders. Little is known about the posttranscriptional regulation of RGS4 expression. We cloned the full-length cDNA of rabbit RGS4, which contains a long 3'-untranslated region (UTR) with several AU-rich elements (AREs). We determined whether RGS4 mRNA stability is mediated by the RNA-binding protein human antigen R (HuR) and contributes to IL-1β-induced upregulation of RGS4 expression. We show that IL-1β treatment in colonic smooth muscle cells doubled the half-life of RGS4 mRNA. Addition of RGS4 3'-UTR to the downstream of Renilla luciferase reporter induced dramatic reduction in the enzyme activity and mRNA expression of luciferase, which was attenuated by the site-directed mutation of the two 3'-most ARE sites. IL-1β increased luciferase mRNA stability in a UTR-dependent manner. Knockdown of HuR significantly aggravated UTR-mediated instability of luciferase and inhibited IL-1β-induced upregulation of RGS4 mRNA. In addition, IL-1β increased cytosolic translocation and RGS4 mRNA binding of HuR. These findings suggest that 3'-most ARE sites within RGS4 3'-UTR are essential for the instability of RGS4 mRNA and IL-1β promotes the stability of RGS4 mRNA through HuR.
Authors:
Fang Li; Danielle Y Hu; Shu Liu; Sunila Mahavadi; William Yen; Karnam S Murthy; Kamel Khalili; Wenhui Hu
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Publication Detail:
Type:  Journal Article; Research Support, N.I.H., Extramural     Date:  2010-09-29
Journal Detail:
Title:  American journal of physiology. Cell physiology     Volume:  299     ISSN:  1522-1563     ISO Abbreviation:  Am. J. Physiol., Cell Physiol.     Publication Date:  2010 Dec 
Date Detail:
Created Date:  2010-12-01     Completed Date:  2011-01-04     Revised Date:  2011-12-21    
Medline Journal Info:
Nlm Unique ID:  100901225     Medline TA:  Am J Physiol Cell Physiol     Country:  United States    
Other Details:
Languages:  eng     Pagination:  C1418-29     Citation Subset:  IM    
Affiliation:
Department of Neuroscience, Temple University School of Medicine, Philadelphia, Pennsylvania 19140, USA.
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MeSH Terms
Descriptor/Qualifier:
3' Untranslated Regions
Animals
Antigens, Surface / genetics,  metabolism*
Base Sequence
Colon / metabolism*
Humans
Interleukin-1beta / metabolism*,  pharmacology
Molecular Sequence Data
Myocytes, Smooth Muscle / metabolism*
RGS Proteins / genetics,  metabolism*
RNA Stability*
RNA-Binding Proteins / genetics,  metabolism*
Rabbits
Up-Regulation
Grant Support
ID/Acronym/Agency:
DK-015564/DK/NIDDK NIH HHS; DK-075964/DK/NIDDK NIH HHS
Chemical
Reg. No./Substance:
0/3' Untranslated Regions; 0/Antigens, Surface; 0/ELAVL1 protein, human; 0/Interleukin-1beta; 0/RGS Proteins; 0/RNA-Binding Proteins; 175335-35-0/RGS4 protein

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