Document Detail


RACK1: a novel substrate for the Src protein-tyrosine kinase.
MedLine Citation:
PMID:  12400005     Owner:  NLM     Status:  MEDLINE    
Abstract/OtherAbstract:
RACK1 is one of a group of PKC-interacting proteins collectively called RACKs (Receptors for Activated C-Kinases). Previously, we showed that RACK1 also interacts with the Src tyrosine kinase, and is an inhibitor of Src activity and cell growth. PKC activation induces the intracellular movement and co-localization of RACK1 and Src, and the tyrosine phosphorylation of RACK1. To determine whether RACK1 is a Src substrate, we assessed phosphorylation of RACK1 by various tyrosine kinases in vitro, and by kinase-active and inactive mutants of Src in vivo. We found that RACK1 is a Src substrate. Moreover, Src activity is necessary for both the tyrosine phosphorylation of RACK1 and the binding of RACK1 to Src's SH2 domain that occur following PKC activation. To identify the tyrosine(s) on RACK1 that is phosphorylated by Src, we generated and tested a series of RACK1 mutants. We found that Src phosphorylates RACK1 on Tyr 228 and/or Tyr 246, highly-conserved tyrosines located in the sixth WD repeat that interact with Src's SH2 domain. We think that RACK1 is an important Src substrate that signals downstream of growth factor receptor tyrosine kinases and is involved in the regulation of Src function and cell growth.
Authors:
Betty Y Chang; Rachel A Harte; Christine A Cartwright
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Publication Detail:
Type:  Journal Article; Research Support, U.S. Gov't, P.H.S.    
Journal Detail:
Title:  Oncogene     Volume:  21     ISSN:  0950-9232     ISO Abbreviation:  Oncogene     Publication Date:  2002 Oct 
Date Detail:
Created Date:  2002-10-25     Completed Date:  2002-11-19     Revised Date:  2009-11-19    
Medline Journal Info:
Nlm Unique ID:  8711562     Medline TA:  Oncogene     Country:  England    
Other Details:
Languages:  eng     Pagination:  7619-29     Citation Subset:  IM    
Affiliation:
Department of Medicine, Stanford University, Stanford, California, CA 94305, USA.
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MeSH Terms
Descriptor/Qualifier:
3T3 Cells
Amino Acid Sequence
Amino Acid Substitution
Animals
Binding Sites
CHO Cells
Cricetinae
Mice
Molecular Sequence Data
Oncogene Proteins v-abl / metabolism
Phosphorylation
Protein Kinase C / metabolism
Receptor, Epidermal Growth Factor / metabolism
Receptors, Cell Surface / genetics,  metabolism*
Receptors, Platelet-Derived Growth Factor / metabolism
Repetitive Sequences, Amino Acid
Sequence Homology, Amino Acid
Substrate Specificity
Tyrosine / metabolism
src Homology Domains
src-Family Kinases / metabolism*
Grant Support
ID/Acronym/Agency:
CA69810/CA/NCI NIH HHS; R01 DK43743/DK/NIDDK NIH HHS
Chemical
Reg. No./Substance:
0/Oncogene Proteins v-abl; 0/Receptors, Cell Surface; 0/receptors for activated C kinase; 55520-40-6/Tyrosine; EC 2.7.10.1/Receptor, Epidermal Growth Factor; EC 2.7.10.1/Receptors, Platelet-Derived Growth Factor; EC 2.7.10.2/src-Family Kinases; EC 2.7.11.13/Protein Kinase C
Comments/Corrections
Erratum In:
Oncogene. 2003 Mar 20;22(11):1748

From MEDLINE®/PubMed®, a database of the U.S. National Library of Medicine


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