Document Detail


Properties of deactivation gating currents in Shaker channels.
MedLine Citation:
PMID:  21354387     Owner:  NLM     Status:  MEDLINE    
Abstract/OtherAbstract:
The charge versus voltage relation of voltage-sensor domains shifts in the voltage axis depending on the initial voltage. Here we show that in nonconducting W434F Shaker K(+) channels, a large portion of this charge-voltage shift is apparent due to a dramatic slowing of the deactivation gating currents, Ig(D) (with τ up to 80 ms), which develops with a time course of ∼1.8 s. This slowing in Ig(D) adds up to the slowing due to pore opening and is absent in the presence of 4-aminopyridine, a compound that prevents the last gating step that leads to pore opening. A remaining 10-15 mV negative shift in the voltage dependence of both the kinetics and the charge movement persists independently of the depolarizing prepulse duration and remains in the presence of 4-aminopyridine, suggesting the existence of an intrinsic offset in the local electric field seen by activated channels. We propose a new (to our knowledge) kinetic model that accounts for these observations.
Authors:
Jérôme J Lacroix; Alain J Labro; Francisco Bezanilla
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Publication Detail:
Type:  Journal Article; Research Support, N.I.H., Extramural    
Journal Detail:
Title:  Biophysical journal     Volume:  100     ISSN:  1542-0086     ISO Abbreviation:  Biophys. J.     Publication Date:  2011 Mar 
Date Detail:
Created Date:  2011-02-28     Completed Date:  2011-06-02     Revised Date:  2012-09-24    
Medline Journal Info:
Nlm Unique ID:  0370626     Medline TA:  Biophys J     Country:  United States    
Other Details:
Languages:  eng     Pagination:  L28-30     Citation Subset:  IM    
Copyright Information:
Copyright © 2011 Biophysical Society. Published by Elsevier Inc. All rights reserved.
Affiliation:
Department of Biochemistry and Molecular Biology, University of Chicago, Chicago, Illinois, USA.
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MeSH Terms
Descriptor/Qualifier:
4-Aminopyridine / pharmacology
Animals
Ion Channel Gating / drug effects,  physiology*
Kinetics
Models, Biological*
Shaker Superfamily of Potassium Channels / metabolism*
Grant Support
ID/Acronym/Agency:
GM030376/GM/NIGMS NIH HHS
Chemical
Reg. No./Substance:
0/Shaker Superfamily of Potassium Channels; 504-24-5/4-Aminopyridine
Comments/Corrections
Comment In:
Biophys J. 2011 Jul 20;101(2):512-4; discussion 515-6   [PMID:  21767505 ]

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