Document Detail


Profiling the array of Ca(v)3.1 variants from the human T-type calcium channel gene CACNA1G: alternative structures, developmental expression, and biophysical variations.
MedLine Citation:
PMID:  16671074     Owner:  NLM     Status:  MEDLINE    
Abstract/OtherAbstract:
We describe the regulated transcriptome of CACNA1G, a human gene for T-type Ca(v)3.1 calcium channels that is subject to extensive alternative RNA splicing. Fifteen sites of transcript variation include 2 alternative 5'-UTR promoter sites, 2 alternative 3'-UTR polyadenylation sites, and 11 sites of alternative splicing within the open reading frame. A survey of 1580 fetal and adult human brain full-length complementary DNAs reveals a family of 30 distinct transcripts, including multiple functional forms that vary in expression with development. Statistical analyses of fetal and adult transcript populations reveal patterns of linkages among intramolecular splice site configurations that change dramatically with development. A shift from nearly independent, biased splicing in fetal transcripts to strongly concerted splicing in adult transcripts suggests progressive activation of multiple "programs" of splicing regulation that reorganize molecular structures in differentiating cells. Patch-clamp studies of nine selected variants help relate splicing regulation to permutations of the gating parameters most likely to modify T-channel physiology in expressing neurons. Gating behavior reflects combinatorial interactions between variable domains so that molecular phenotype depends on ensembles of coselected domains, consistent with the observed emergence of concerted splicing during development. We conclude that the structural gene and networks of splicing regulatory factors define an integrated system for the phenotypic variation of Ca(v)3.1 biophysics during nervous system development.
Authors:
Mark C Emerick; Rebecca Stein; Robin Kunze; Megan M McNulty; Melissa R Regan; Dorothy A Hanck; William S Agnew
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Publication Detail:
Type:  Journal Article; Research Support, N.I.H., Extramural; Research Support, Non-U.S. Gov't    
Journal Detail:
Title:  Proteins     Volume:  64     ISSN:  1097-0134     ISO Abbreviation:  Proteins     Publication Date:  2006 Aug 
Date Detail:
Created Date:  2006-06-22     Completed Date:  2006-08-02     Revised Date:  2008-11-21    
Medline Journal Info:
Nlm Unique ID:  8700181     Medline TA:  Proteins     Country:  United States    
Other Details:
Languages:  eng     Pagination:  320-42     Citation Subset:  IM    
Copyright Information:
Copyright 2006 Wiley-Liss, Inc.
Affiliation:
Department of Physiology, Johns Hopkins University School of Medicine, Baltimore, Maryland 21205, USA.
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MeSH Terms
Descriptor/Qualifier:
Alternative Splicing
Biophysics / methods
Brain / embryology,  metabolism
Calcium Channels, T-Type / chemistry,  physiology*
DNA, Complementary / metabolism
Gene Expression Regulation, Developmental*
Genetic Variation
Humans
Kinetics
Open Reading Frames
Patch-Clamp Techniques
Protein Conformation
Protein Structure, Tertiary
Grant Support
ID/Acronym/Agency:
HL-R01-HL62603/HL/NHLBI NIH HHS
Chemical
Reg. No./Substance:
0/CACNA1G protein, human; 0/Calcium Channels, T-Type; 0/DNA, Complementary

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