Document Detail


Pharmacological inhibition of neuronal NADPH oxidase protects against 1-methyl-4-phenylpyridinium (MPP+)-induced oxidative stress and apoptosis in mesencephalic dopaminergic neuronal cells.
MedLine Citation:
PMID:  17904225     Owner:  NLM     Status:  MEDLINE    
Abstract/OtherAbstract:
Oxidative stress is widely recognized as a key mediator of degenerative processes in Parkinson's disease (PD). Recently, we demonstrated that the dopaminergic toxin MPP+ initiates oxidative stress to cause caspase-3-dependent apoptotic cell death in mesencephalic dopaminergic neuronal (N27) cells. In this study, we determined the source of reactive oxygen species (ROS) produced during MPP+-induced apoptotic cell death. In addition to mitochondria, plasma membrane NADPH oxidase is considered a major producer of ROS inside the cell. Here, we show that N27 neuronal cells express key NADPH oxidase subunits gp91phox and p67phox. We used structurally diverse NADPH oxidase inhibitors, aminoethyl-benzenesulfonylfluoride (AEBSF, 100-1000microM), apocynin (100-1000microM), and diphenylene iodonium (DPI, 3-30microM), to inhibit intrinsic NADPH oxidase activity in N27 cells. Flow cytometric analysis using the ROS-sensitive dye hydroethidine revealed that AEBSF blocked 300microM MPP+-induced ROS production for over 45min in N27 cells, in a dose-dependent manner. Further treatment with DPI, apocynin, and SOD also blocked MPP+-induced ROS production. In Sytox cell death assays, co-treatment with AEBSF, apocynin, or DPI for 24h significantly suppressed MPP+-induced cytotoxic cell death. Similarly, co-treatment with these inhibitors also significantly attenuated MPP+-induced increases in caspase-3 enzymatic activity. Furthermore, quantitative DNA fragmentation ELISA assays revealed that AEBSF, DPI, and apocynin rescue N27 cells from MPP+-induced apoptotic cell death. Together, these results indicate for the first time that intracellular ROS generated by NAPDH oxidase are present within the mesencephalic neuronal cells, and are a key determinant of MPP+-mediated dopaminergic degeneration in in vitro models of dopaminergic degeneration. This study supports a critical role of NADPH oxidase in the oxidative damage in PD; targeting this enzyme may lead to novel therapies for PD.
Authors:
Vellareddy Anantharam; Siddharth Kaul; Chunjuan Song; Arthi Kanthasamy; Anumantha G Kanthasamy
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Publication Detail:
Type:  Journal Article; Research Support, N.I.H., Extramural     Date:  2007-08-25
Journal Detail:
Title:  Neurotoxicology     Volume:  28     ISSN:  0161-813X     ISO Abbreviation:  Neurotoxicology     Publication Date:  2007 Sep 
Date Detail:
Created Date:  2007-10-09     Completed Date:  2007-12-26     Revised Date:  2011-04-26    
Medline Journal Info:
Nlm Unique ID:  7905589     Medline TA:  Neurotoxicology     Country:  Netherlands    
Other Details:
Languages:  eng     Pagination:  988-97     Citation Subset:  IM    
Affiliation:
Parkinson's Disorder Research Laboratory, Iowa Center for Advanced Neurotoxicology, Department of Biomedical Sciences, Iowa State University, Ames, IA 50011-1250, USA.
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MeSH Terms
Descriptor/Qualifier:
1-Methyl-4-phenylpyridinium / antagonists & inhibitors*,  toxicity*
Animals
Apoptosis / drug effects*
Blotting, Western
Caspases / metabolism
Cell Survival / drug effects
DNA Fragmentation / drug effects
Dopamine / physiology*
Dopamine Agents / toxicity*
Dose-Response Relationship, Drug
Enzyme Inhibitors / pharmacology*
MPTP Poisoning / metabolism*,  prevention & control*
Membrane Glycoproteins / genetics
Mesencephalon / cytology,  drug effects*
NADPH Oxidase / antagonists & inhibitors*,  genetics
Neurons / drug effects,  enzymology*
Organic Chemicals
Oxidative Stress / drug effects*
Phosphoproteins / genetics
Rats
Reactive Oxygen Species / metabolism
Grant Support
ID/Acronym/Agency:
ES10586/ES/NIEHS NIH HHS; NS38644/NS/NINDS NIH HHS; NS45133/NS/NINDS NIH HHS; R01 ES010586-07/ES/NIEHS NIH HHS; R01 NS038644-09/NS/NINDS NIH HHS
Chemical
Reg. No./Substance:
0/Dopamine Agents; 0/Enzyme Inhibitors; 0/Membrane Glycoproteins; 0/Organic Chemicals; 0/Phosphoproteins; 0/Reactive Oxygen Species; 0/SYTOX Green; 0/neutrophil cytosol factor 67K; 48134-75-4/1-Methyl-4-phenylpyridinium; EC 1.6.-/Cybb protein, rat; EC 1.6.3.1/NADPH Oxidase; EC 3.4.22.-/Caspases
Comments/Corrections

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