Document Detail


PapX, a P fimbrial operon-encoded inhibitor of motility in uropathogenic Escherichia coli.
MedLine Citation:
PMID:  18710869     Owner:  NLM     Status:  MEDLINE    
Abstract/OtherAbstract:
Motility and adherence are two integral aspects of bacterial pathogenesis. Adherence, often mediated by fimbriae, permits bacteria to attach to host cells and establish infection, whereas flagellum-driven motility allows bacteria to disseminate to sites more advantageous for colonization. Both fimbriae and flagella have been proven important for virulence of uropathogenic Escherichia coli (UPEC). Reciprocal regulation is one mechanism by which bacteria may reconcile the contradictory actions of adherence and motility. PapX, a P fimbrial gene product of UPEC strain CFT073, is a functional homolog of MrpJ of Proteus mirabilis; ectopic expression of papX in P. mirabilis reduces motility. To define the connection between P fimbria expression and motility in UPEC, the role of papX in the regulation of motility of strain CFT073 was examined. Overexpression of papX decreased motility of CFT073, which correlated with both a significant reduction in flagellin protein synthesized and flagella assembled on the cell surface. Conversely, an increase in motility and flagellin production was seen in an isogenic papX deletion mutant of CFT073. Microarray and quantitative reverse transcription-PCR analysis indicated that repression of motility of CFT073 by PapX appears to occur at the transcriptional level; expression of many motility-associated genes, including flhDC, the master regulator of motility, is decreased when papX is overexpressed. Transcription of motility genes is increased in the papX mutant compared to wild type. Electrophoretic mobility gel shift analysis revealed that PapX binds to the flhD promoter. We conclude that synthesis of P fimbriae regulates flagellum synthesis to repress motility via PapX.
Authors:
Amy N Simms; Harry L T Mobley
Related Documents :
12864849 - Complex regulation of csgd promoter activity by global regulatory proteins.
9268019 - Transcriptional regulation of pilc2 in neisseria gonorrhoeae: response to oxygen availa...
20639319 - Posttranscriptional regulation of glutamine synthetase in the filamentous cyanobacteriu...
20656909 - The novel two-component regulatory system bfisr regulates biofilm development by contro...
16468999 - Pspb and pspc of yersinia enterocolitica are dual function proteins: regulators and eff...
12576599 - Differential expression of candida albicans phospholipase b (plb1) under various enviro...
12864849 - Complex regulation of csgd promoter activity by global regulatory proteins.
9490789 - Nf-kappab mediated transcriptional activation is enhanced by the architectural factor h...
11306579 - Hsp-cbf is an nf-y-dependent coactivator of the heat shock promoters ccaat boxes.
Publication Detail:
Type:  Journal Article; Research Support, N.I.H., Extramural     Date:  2008-08-18
Journal Detail:
Title:  Infection and immunity     Volume:  76     ISSN:  1098-5522     ISO Abbreviation:  Infect. Immun.     Publication Date:  2008 Nov 
Date Detail:
Created Date:  2008-10-20     Completed Date:  2008-11-18     Revised Date:  2009-11-18    
Medline Journal Info:
Nlm Unique ID:  0246127     Medline TA:  Infect Immun     Country:  United States    
Other Details:
Languages:  eng     Pagination:  4833-41     Citation Subset:  IM    
Affiliation:
Department of Microbiology and Immunology, University of Michigan Medical School, Ann Arbor, MI 48109, USA.
Export Citation:
APA/MLA Format     Download EndNote     Download BibTex
MeSH Terms
Descriptor/Qualifier:
Animals
Bacterial Adhesion
Electrophoretic Mobility Shift Assay
Escherichia coli / pathogenicity*
Escherichia coli Proteins / genetics*,  metabolism
Fimbriae Proteins / genetics*
Fimbriae, Bacterial / genetics*,  metabolism
Flagella / genetics,  metabolism
Flagellin / genetics,  metabolism
Fluorescent Antibody Technique, Indirect
Gene Expression
Gene Expression Regulation, Bacterial
Mice
Oligonucleotide Array Sequence Analysis
Operon / genetics
Reverse Transcriptase Polymerase Chain Reaction
Transcription, Genetic
Urinary Tract Infections / microbiology*
Grant Support
ID/Acronym/Agency:
AI059722/AI/NIAID NIH HHS
Chemical
Reg. No./Substance:
0/Escherichia coli Proteins; 12777-81-0/Flagellin; 147680-16-8/Fimbriae Proteins
Comments/Corrections

From MEDLINE®/PubMed®, a database of the U.S. National Library of Medicine


Previous Document:  ADP-ribosylation of actin by the Clostridium botulinum C2 toxin in mammalian cells results in delaye...
Next Document:  Total, membrane, and immunogenic proteomes of macrophage- and tick cell-derived Ehrlichia chaffeensi...