Document Detail


Nucleotide excision by E. coli DNA polymerase I in proofreading and non-proofreading modes.
MedLine Citation:
PMID:  3061468     Owner:  NLM     Status:  MEDLINE    
Abstract/OtherAbstract:
Escherichia coli DNA polymerase I exists in at least two distinct kinetic forms. When it binds to a template, the proofreading activity is usually switched off. As the enzyme progresses along the template, it becomes more and more competent for excision. This phenomenon introduces a link between fidelity and processivity. Processivity is best studied when the chain-length distributions of synthesized polymers are stationary. Even then, however, one cannot avoid multiple initiations on a given template by the same molecule of the enzyme. When synthesis is initiated with primers of lengths 15 or 20, a strange phenomenon is observed. It seems that the polymerase starts by hydrolyzing the primer down to a length of 7-10 nucleotides and only then starts to add nucleotides. It does so in a low-accuracy mode, suggesting that, while the exonuclease is clearly active, it does not contribute to proofreading. The warm-up of the proofreading function is therefore reinterpreted as a switch between two modes of behaviour: a mode 1 of low accuracy in which the 3'----5' exonuclease, while active, is uncoupled from the polymerase and does not contribute to proofreading, and a mode 2 of high accuracy in which the exonuclease is kinetically linked to the polymerase activity.
Authors:
P J Lecomte; J Ninio
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Publication Detail:
Type:  Journal Article; Research Support, Non-U.S. Gov't    
Journal Detail:
Title:  Biochimica et biophysica acta     Volume:  951     ISSN:  0006-3002     ISO Abbreviation:  Biochim. Biophys. Acta     Publication Date:  1988 Dec 
Date Detail:
Created Date:  1989-02-13     Completed Date:  1989-02-13     Revised Date:  2006-11-15    
Medline Journal Info:
Nlm Unique ID:  0217513     Medline TA:  Biochim Biophys Acta     Country:  NETHERLANDS    
Other Details:
Languages:  eng     Pagination:  255-60     Citation Subset:  IM    
Affiliation:
Institut Jacques Monod, Paris, France.
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MeSH Terms
Descriptor/Qualifier:
DNA Polymerase I / metabolism*
DNA, Bacterial / metabolism*
Escherichia coli / enzymology*,  genetics
Exodeoxyribonuclease V
Exodeoxyribonucleases / metabolism
Kinetics
Nucleotides / metabolism*
Oligodeoxyribonucleotides / metabolism
Poly A / metabolism
Templates, Genetic
Chemical
Reg. No./Substance:
0/DNA, Bacterial; 0/Nucleotides; 0/Oligodeoxyribonucleotides; 0/oligo (dT); 24937-83-5/Poly A; 25191-20-2/poly(dA); EC 2.7.7.-/DNA Polymerase I; EC 3.1.-/Exodeoxyribonucleases; EC 3.1.11.5/Exodeoxyribonuclease V

From MEDLINE®/PubMed®, a database of the U.S. National Library of Medicine


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