Document Detail


Nuclear retention of fission yeast dicer is a prerequisite for RNAi-mediated heterochromatin assembly.
MedLine Citation:
PMID:  20152181     Owner:  NLM     Status:  MEDLINE    
Abstract/OtherAbstract:
RNaseIII ribonucleases act at the heart of RNA silencing pathways by processing precursor RNAs into mature microRNAs and siRNAs. In the fission yeast Schizosaccharomyces pombe, siRNAs are generated by the RNaseIII enzyme Dcr1 and are required for heterochromatin formation at centromeres. In this study, we have analyzed the subcellular localization of Dcr1 and found that it accumulates in the nucleus and is enriched at the nuclear periphery. Nuclear accumulation of Dcr1 depends on a short motif that impedes nuclear export promoted by the double-stranded RNA binding domain of Dcr1. Absence of this motif renders Dcr1 mainly cytoplasmic and is accompanied by remarkable changes in gene expression and failure to assemble heterochromatin. Our findings suggest that Dicer proteins are shuttling proteins and that the steady-state subcellular levels can be shifted toward either compartment.
Authors:
Stephan Emmerth; Heiko Schober; Dimos Gaidatzis; Tim Roloff; Kirsten Jacobeit; Marc B?hler
Publication Detail:
Type:  Journal Article; Research Support, Non-U.S. Gov't    
Journal Detail:
Title:  Developmental cell     Volume:  18     ISSN:  1878-1551     ISO Abbreviation:  Dev. Cell     Publication Date:  2010 Jan 
Date Detail:
Created Date:  2010-02-15     Completed Date:  2010-03-19     Revised Date:  -    
Medline Journal Info:
Nlm Unique ID:  101120028     Medline TA:  Dev Cell     Country:  United States    
Other Details:
Languages:  eng     Pagination:  102-13     Citation Subset:  IM    
Copyright Information:
(c) 2010 Elsevier Inc. All rights reserved.
Affiliation:
Friedrich Miescher Institute for Biomedical Research, Maulbeerstrasse 66, CH-4058 Basel, Switzerland.
Data Bank Information
Bank Name/Acc. No.:
GEO/GSE18440;  GSE18582
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MeSH Terms
Descriptor/Qualifier:
Active Transport, Cell Nucleus / genetics
Amino Acid Motifs / genetics
Binding Sites / genetics
Cell Compartmentation / genetics
Cell Nucleus / enzymology,  genetics*
Centromere / genetics,  metabolism,  ultrastructure
Cytoplasm / enzymology,  genetics
Gene Expression Regulation, Fungal / genetics
Heterochromatin / genetics*
Protein Transport / genetics
RNA Interference / physiology*
RNA, Small Interfering / biosynthesis,  genetics
Ribonuclease III / genetics,  metabolism*
Schizosaccharomyces / enzymology,  genetics*
Chemical
Reg. No./Substance:
0/Heterochromatin; 0/RNA, Small Interfering; EC 3.1.26.3/Ribonuclease III
Comments/Corrections
Comment In:
Dev Cell. 2010 Jan 19;18(1):6-7   [PMID:  20152173 ]

From MEDLINE®/PubMed®, a database of the U.S. National Library of Medicine


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