Document Detail


Molecular mechanism of the reduction of cysteine sulfinic acid of peroxiredoxin to cysteine by mammalian sulfiredoxin.
MedLine Citation:
PMID:  16565085     Owner:  NLM     Status:  MEDLINE    
Abstract/OtherAbstract:
Among many proteins with cysteine sulfinic acid (Cys-SO2H) residues, the sulfinic forms of certain peroxiredoxins (Prxs) are selectively reduced by sulfiredoxin (Srx) in the presence of ATP. All Srx enzymes contain a conserved cysteine residue. To elucidate the mechanism of the Srx-catalyzed reaction, we generated various mutants of Srx and examined their interaction with PrxI, their ATPase activity, and their ability to reduce sulfinic PrxI. Our results suggest that three surface-exposed amino acid residues, corresponding to Arg50, Asp57, and Asp79 of rat Srx, are critical for substrate recognition. The presence of the sulfinic form (but not the reduced form) of PrxI induces the conserved cysteine of Srx to take the gamma-phosphate of ATP and then immediately transfers the phosphate to the sulfinic moiety of PrxI to generate a sulfinic acid phosphoryl ester (Prx-Cys-S(=O)OPO3(2-)). This ester is reductively cleaved by a thiol molecule (RSH) such as GSH, thioredoxin, and dithiothreitol to produce a disulfide-S-monoxide (Prx-Cys-S(=O)-S-R). The disulfide-S-monoxide is further reduced through the oxidation of three thiol equivalents to complete the catalytic cycle and regenerate Prx-Cys-SH.
Authors:
Woojin Jeong; Sung Jun Park; Tong-Shin Chang; Duck-Yeon Lee; Sue Goo Rhee
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Publication Detail:
Type:  Journal Article; Research Support, Non-U.S. Gov't     Date:  2006-03-24
Journal Detail:
Title:  The Journal of biological chemistry     Volume:  281     ISSN:  0021-9258     ISO Abbreviation:  J. Biol. Chem.     Publication Date:  2006 May 
Date Detail:
Created Date:  2006-05-15     Completed Date:  2006-07-21     Revised Date:  2009-11-19    
Medline Journal Info:
Nlm Unique ID:  2985121R     Medline TA:  J Biol Chem     Country:  United States    
Other Details:
Languages:  eng     Pagination:  14400-7     Citation Subset:  IM    
Affiliation:
Laboratory of Cell Signaling, National Heart, Lung, and Blood Institute, National Institutes of Health, Bethesda, Maryland 20892, USA.
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MeSH Terms
Descriptor/Qualifier:
Adenosine Diphosphate / chemistry
Adenosine Triphosphate / chemistry
Animals
Catalysis
Cysteine / analogs & derivatives*,  chemistry*
Disulfides / chemistry
Dithiothreitol / chemistry
Humans
Hydrolysis
Models, Chemical
Models, Molecular
Mutation
Oxidoreductases / chemistry*
Oxidoreductases Acting on Sulfur Group Donors
Peroxidases / chemistry*
Peroxiredoxins
Phosphates / chemistry
Phosphorylation
Rats
Chemical
Reg. No./Substance:
0/Disulfides; 0/Phosphates; 2381-08-0/cysteine sulfinic acid; 3483-12-3/Dithiothreitol; 52-90-4/Cysteine; 56-65-5/Adenosine Triphosphate; 58-64-0/Adenosine Diphosphate; EC 1.-/Oxidoreductases; EC 1.11.1.-/Peroxidases; EC 1.11.1.15/Peroxiredoxins; EC 1.8.-/Oxidoreductases Acting on Sulfur Group Donors; EC 1.8.98.2/SRXN1 protein, human

From MEDLINE®/PubMed®, a database of the U.S. National Library of Medicine


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