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Molecular Mechanism of Interleukin-2-induced Mucosal Homeostasis.
MedLine Citation:
PMID:  22116305     Owner:  NLM     Status:  Publisher    
Abstract/OtherAbstract:
Sustained damage to the mucosal lining in patients with Inflammatory Bowel Disease (IBD) facilitates translocation of intestinal microbes to sub-mucosal immune cells leading to chronic inflammation. Previously we demonstrated the role of Jak3 in IL-2-induced IEC migration, one of the early events during intestinal wound repair. In this report we demonstrate that IL-2 also plays a role in IEC homeostasis through concentration-dependent regulation of IEC proliferation and cell-death. At lower concentrations (≤ 50U/ml) IL-2 promoted proliferation, while at higher concentrations (100 U/ml), it promoted apoptosis. Activation by IL-2 led to tyrosine phosphorylation-dependent interactions between Jak3 and p52ShcA only at lower concentrations. Phosphatase SHP1 dephosphorylated IL-2-induced phosphorylated p52ShcA. Higher concentrations of IL-2 decreased the phosphorylation of Jak3 and p52ShcA, disrupted their interactions, redistributed Jak3 to the nucleus, and induced apoptosis in IEC. IL-2 also induced dose dependent up-regulation of p52shcA, and down-regulation of jak3-mRNA. Constitutive over-expression and mir-shRNA mediated knock-down studies showed that expression of both Jak3 and p52ShcA were necessary for IL-2-induced proliferation of IEC. Doxycycline regulated sh-RNA-expression demonstrated that IL-2-induced down regulation of jak3-mRNA was responsible for higher IL-2-induced apoptosis in IEC. Collectively, these data demonstrate a novel mechanism of IL-2-induced mucosal homeostasis through post-translational and transcriptional regulation of Jak3 and p52ShcA.
Authors:
Jayshree Mishra; Christopher M Waters; Narendra Kumar
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Publication Detail:
Type:  JOURNAL ARTICLE     Date:  2011-11-23
Journal Detail:
Title:  American journal of physiology. Cell physiology     Volume:  -     ISSN:  1522-1563     ISO Abbreviation:  -     Publication Date:  2011 Nov 
Date Detail:
Created Date:  2011-11-25     Completed Date:  -     Revised Date:  -    
Medline Journal Info:
Nlm Unique ID:  100901225     Medline TA:  Am J Physiol Cell Physiol     Country:  -    
Other Details:
Languages:  ENG     Pagination:  -     Citation Subset:  -    
Affiliation:
Texas A&M Health Sciences Center.
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