Document Detail


Modelling the peroxisomal carbon leak during lipid mobilization in Arabidopsis.
MedLine Citation:
PMID:  20863290     Owner:  NLM     Status:  MEDLINE    
Abstract/OtherAbstract:
Mutation of the ACN1 (acetate non-utilizing 1) locus of Arabidopsis results in altered acetate assimilation into gluconeogenic sugars and anapleurotic amino acids and leads to an overall depression in primary metabolite levels by approx. 50% during seedling development. Levels of acetyl-CoA were higher in acn1 compared with wild-type, which is counterintuitive to the activity of ACN1 as a peroxisomal acetyl-CoA synthetase. We hypothesize that ACN1 recycles free acetate to acetyl-CoA within peroxisomes in order that carbon remains fed into the glyoxylate cycle. When ACN1 is not present, carbon in the form of acetate can leak out of peroxisomes and is reactivated to acetyl-CoA within the cytosol. Kinetic models incorporating estimates of carbon input and pathway dynamics from a variety of literature sources have proven useful in explaining how ACN1 may prevent the carbon leak and even contribute to the control of peroxisomal carbon metabolism.
Authors:
Mark A Hooks; Elizabeth Allen; Jonathan A D Wattis
Publication Detail:
Type:  Journal Article; Research Support, Non-U.S. Gov't    
Journal Detail:
Title:  Biochemical Society transactions     Volume:  38     ISSN:  1470-8752     ISO Abbreviation:  Biochem. Soc. Trans.     Publication Date:  2010 Oct 
Date Detail:
Created Date:  2010-09-24     Completed Date:  2011-01-06     Revised Date:  -    
Medline Journal Info:
Nlm Unique ID:  7506897     Medline TA:  Biochem Soc Trans     Country:  England    
Other Details:
Languages:  eng     Pagination:  1230-3     Citation Subset:  IM    
Affiliation:
School of Biological Sciences, Bangor University, Bangor, Gwynedd LL57 2UW, UK. m.a.hooks@bangor.ac.uk
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MeSH Terms
Descriptor/Qualifier:
Acetates / metabolism
Acetyl Coenzyme A / metabolism
Arabidopsis / genetics,  metabolism*
Arabidopsis Proteins / genetics,  metabolism
Carbon / metabolism*
Coenzyme A Ligases / genetics,  metabolism
Lipid Metabolism
Peroxisomes / metabolism*
Grant Support
ID/Acronym/Agency:
P14659//Biotechnology and Biological Sciences Research Council; P19408//Biotechnology and Biological Sciences Research Council
Chemical
Reg. No./Substance:
0/Acetates; 0/Arabidopsis Proteins; 72-89-9/Acetyl Coenzyme A; 7440-44-0/Carbon; EC 6.2.1.-/Coenzyme A Ligases; EC 6.2.1.13/acetate-CoA ligase (ADP-forming)

From MEDLINE®/PubMed®, a database of the U.S. National Library of Medicine


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