Document Detail


Modeling light-induced currents in the eye of Chlamydomonas reinhardtii.
MedLine Citation:
PMID:  12235485     Owner:  NLM     Status:  MEDLINE    
Abstract/OtherAbstract:
Rhodopsin-mediated electrical events in green algae have been recorded in the past from the eyes of numerous micro-algae like Haematococcus pluvialis, Chlamydomonas reinhardtii and Volvox carteri. However, the electrical data gathered by suction-pipette techniques could be interpreted in qualitative terms only. Here we present two models that allow a quantitative analysis of such results: First, an electrical analog circuit for the cell in suction pipette configuration is established. Applying this model to experimental data from unilluminated cells of C. reinhardtii yields a membrane conductance of about 3 Sm(-2). Furthermore, an analog circuit allows the determination of the photocurrent fraction that is recorded under experimental conditions. Second, a reaction scheme of a rhodopsin-type photocycle with an early Ca(2+) conductance and a later H(+) conductance is presented. The combination of both models provides good fits to light-induced currents recorded from C. reinhardtii. Finally, it allowed the calculation of the impact of each model parameter on the time courses of observable photocurrent and of inferred transmembrane voltage. The reduction of the flash-to-peak times at increasing light intensities are explained by superposition of two kinetically distinct rhodopsins and by assuming that the Ca(2+)-conducting state decays faster at more positive membrane voltages.
Authors:
D Gradmann; S Ehlenbeck; P Hegemann
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Publication Detail:
Type:  Comparative Study; Journal Article; Research Support, Non-U.S. Gov't    
Journal Detail:
Title:  The Journal of membrane biology     Volume:  189     ISSN:  0022-2631     ISO Abbreviation:  J. Membr. Biol.     Publication Date:  2002 Sep 
Date Detail:
Created Date:  2002-09-17     Completed Date:  2003-04-04     Revised Date:  2008-11-21    
Medline Journal Info:
Nlm Unique ID:  0211301     Medline TA:  J Membr Biol     Country:  United States    
Other Details:
Languages:  eng     Pagination:  93-104     Citation Subset:  IM    
Affiliation:
Abteilung Biophysik der Pflanze, Albrecht-von-Haller-Institut für Pflanzenwissenschaften der Universität, Untere Karspüle 2, D-37073 Göttingen, Germany. dgradma@gwdg.de
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MeSH Terms
Descriptor/Qualifier:
Algorithms
Animals
Chlamydomonas reinhardtii / physiology*,  radiation effects*
Computer Simulation
Dose-Response Relationship, Radiation
Electric Conductivity
Light*
Membrane Potentials / physiology,  radiation effects
Models, Biological*
Photobiology / methods
Photoperiod
Photoreceptor Cells, Invertebrate / physiology*,  radiation effects*
Rhodopsins, Microbial / physiology*
Sensitivity and Specificity
Chemical
Reg. No./Substance:
0/Rhodopsins, Microbial

From MEDLINE®/PubMed®, a database of the U.S. National Library of Medicine


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