Document Detail


MT1-MMP is required for myeloid cell fusion via regulation of Rac1 signaling.
MedLine Citation:
PMID:  20152179     Owner:  NLM     Status:  MEDLINE    
Abstract/OtherAbstract:
Cell fusion is essential for fertilization, myotube formation, and inflammation. Macrophages fuse under various circumstances, but the molecular signals involved in the distinct steps of their fusion are not fully characterized. Using null mice and derived cells, we show that the protease MT1-MMP is necessary for macrophage fusion during osteoclast and giant-cell formation in vitro and in vivo. Specifically, MT1-MMP is required for lamellipodia formation and for proper cell morphology and motility of bone marrow myeloid progenitors prior to membrane fusion. These functions of MT1-MMP do not depend on MT1-MMP catalytic activity or downstream pro-MMP-2 activation. Instead, MT1-MMP null cells show a decreased Rac1 activity and reduced membrane targeting of Rac1 and the adaptor protein p130Cas. Retroviral rescue experiments and protein binding assays delineate a signaling pathway in which MT1-MMP, via its cytosolic tail, contributes to macrophage migration and fusion by regulating Rac1 activity through an association with p130Cas.
Authors:
Pilar Gonzalo; Marta C Guadamillas; María Victoria Hernández-Riquer; Angela Pollán; Araceli Grande-García; Rubén A Bartolomé; Amit Vasanji; Chiara Ambrogio; Roberto Chiarle; Joaquín Teixidó; Juha Risteli; Suneel S Apte; Miguel A del Pozo; Alicia G Arroyo
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Publication Detail:
Type:  Journal Article; Research Support, N.I.H., Extramural; Research Support, Non-U.S. Gov't    
Journal Detail:
Title:  Developmental cell     Volume:  18     ISSN:  1878-1551     ISO Abbreviation:  Dev. Cell     Publication Date:  2010 Jan 
Date Detail:
Created Date:  2010-02-15     Completed Date:  2010-03-19     Revised Date:  2011-07-20    
Medline Journal Info:
Nlm Unique ID:  101120028     Medline TA:  Dev Cell     Country:  United States    
Other Details:
Languages:  eng     Pagination:  77-89     Citation Subset:  IM    
Copyright Information:
(c) 2010 Elsevier Inc. All rights reserved.
Affiliation:
Department of Vascular Biology and Inflammation, Centro Nacional de Investigaciones Cardiovasculares, 28029 Madrid, Spain.
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MeSH Terms
Descriptor/Qualifier:
Animals
Bone Remodeling / physiology
Cell Differentiation / physiology
Cell Fusion
Cell Membrane / metabolism,  ultrastructure
Cell Movement / physiology
Cell Shape / physiology
Cells, Cultured
Crk-Associated Substrate Protein / metabolism
Giant Cells / metabolism,  ultrastructure
Matrix Metalloproteinase 14 / genetics,  metabolism*
Mice
Mice, Inbred C57BL
Mice, Knockout
Myeloid Cells / cytology,  metabolism*
Osteoclasts / cytology,  metabolism*
Protein Structure, Tertiary / physiology
Pseudopodia / metabolism,  ultrastructure
Signal Transduction / physiology
Stem Cells / cytology,  metabolism
rac1 GTP-Binding Protein / genetics,  metabolism*
Grant Support
ID/Acronym/Agency:
AR47074/AR/NIAMS NIH HHS; R01 AR047074-04/AR/NIAMS NIH HHS
Chemical
Reg. No./Substance:
0/Bcar1 protein, mouse; 0/Crk-Associated Substrate Protein; EC 3.4.24.80/Matrix Metalloproteinase 14; EC 3.6.5.2/rac1 GTP-Binding Protein
Comments/Corrections
Comment In:
Dev Cell. 2010 Jan 19;18(1):3-4   [PMID:  20152171 ]

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