Document Detail


Loss of Ikkbeta promotes migration and proliferation of mouse embryo fibroblast cells.
MedLine Citation:
PMID:  16966325     Owner:  NLM     Status:  MEDLINE    
Abstract/OtherAbstract:
The IkappaB kinase complex (IKK) is central to the activation of NF-kappaB, a critical transcription factor governing expression of genes involved in cell proliferation and anti-apoptotic responses. Mice with genetic disruptions of the Ikkbeta or Ikkgamma gene loci die during embryogenesis because of severe hepatic apoptosis. We now show that Ikkbeta gene deficiency promotes migration and proliferation of mouse embryo fibroblast cells. Morphological analyses revealed an unusual protrusion of the cytoplasm in Ikkbeta(-/-) cells when cultured at a lower density. In a Boyden chamber assay, Ikkbeta(-/-) cells exhibited a high rate of invasion and migration. Enhanced formation of actin stress fibers was also observed in the Ikkbeta(-/-) cells. Mechanistic studies indicated that IKKbeta affects the expression of proteins involved in the assembly of cytoskeleton and cell movement. Furthermore, re-expression of Ikkbeta and antioxidant treatment in Ikkbeta(-/-) cells caused a reversal of protrusive phenotype and high motility, respectively. Furthermore, elimination of reactive oxygen species (ROS) blocked expression of snail and subsequently derepressed E-cadherin expression. Although the underlying mechanism is likely entangled and complicated, the data presented indicate that generation of ROS played a key role in the morphological and mobility changes in Ikkbeta(-/-) cells. These data thus suggest that IKKbeta provides inhibitory signals for cell mobility and growth. Deficiency in the Ikkbeta gene promotes cell mobilization, at least partially, through a ROS-dependent mechanism.
Authors:
Fei Chen; Yongju Lu; Vince Castranova; Zhiwei Li; Michael Karin
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Publication Detail:
Type:  Journal Article     Date:  2006-09-11
Journal Detail:
Title:  The Journal of biological chemistry     Volume:  281     ISSN:  0021-9258     ISO Abbreviation:  J. Biol. Chem.     Publication Date:  2006 Dec 
Date Detail:
Created Date:  2006-11-27     Completed Date:  2007-01-30     Revised Date:  2009-11-19    
Medline Journal Info:
Nlm Unique ID:  2985121R     Medline TA:  J Biol Chem     Country:  United States    
Other Details:
Languages:  eng     Pagination:  37142-9     Citation Subset:  IM    
Affiliation:
Health Effects Laboratory Division, National Institute for Occupational Safety and Health, Morgantown, West Virginia 26505, USA. lfd3@cdc.gov
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MeSH Terms
Descriptor/Qualifier:
Animals
Antioxidants / pharmacology
Apoptosis
Cell Adhesion
Cell Movement
Cell Proliferation
Embryo, Mammalian / cytology*
Epithelial Cells / metabolism
Fibroblasts / metabolism*
Gene Expression Regulation*
Humans
I-kappa B Kinase / metabolism,  physiology*
Mice
Phenotype
RNA Interference
Reactive Oxygen Species
Chemical
Reg. No./Substance:
0/Antioxidants; 0/Reactive Oxygen Species; EC 2.7.11.10/I-kappa B Kinase

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