Document Detail


Leishmania donovani amastigotes impair gamma interferon-induced STAT1alpha nuclear translocation by blocking the interaction between STAT1alpha and importin-alpha5.
MedLine Citation:
PMID:  20566692     Owner:  NLM     Status:  MEDLINE    
Abstract/OtherAbstract:
The protozoan parasite Leishmania donovani, the etiological agent of visceral leishmaniasis, is renowned for its capacity to sabotage macrophage functions and signaling pathways stimulated by activators such as gamma interferon (IFN-gamma). Our knowledge of the strategies utilized by L. donovani to impair macrophage responsiveness to IFN-gamma remains fragmentary. In the present study, we investigated the impact of an infection by the amastigote stage of L. donovani on IFN-gamma responses and signaling via the Janus kinase (JAK)-signal transducer and activator of transcription (STAT) pathway in mouse bone marrow-derived macrophages. The levels of IFN-gamma-induced expression of major histocompatibility complex class II and inducible nitric oxide synthase (iNOS) were strongly reduced in L. donovani amastigote-infected macrophages. As the expression of those genes is mediated by the transcription factors STAT1alpha and IFN regulatory factor 1 (IRF-1), we investigated their activation in amastigote-infected macrophages treated with IFN-gamma. We found that whereas STAT1alpha protein levels and the levels of phosphorylation on Tyr701 and Ser727 were normal, IRF-1 expression was inhibited in infected macrophages. This inhibition of IRF-1 expression correlated with a defective nuclear translocation of STAT1alpha, and further analyses revealed that the IFN-gamma-induced STAT1alpha association with the nuclear transport adaptor importin-alpha5 was compromised in L. donovani amastigote-infected macrophages. Taken together, our results provide evidence for a novel mechanism used by L. donovani amastigotes to interfere with IFN-gamma-activated macrophage functions and provide a better understanding of the strategies deployed by this parasite to ensure its intracellular survival.
Authors:
Christine Matte; Albert Descoteaux
Publication Detail:
Type:  Journal Article; Research Support, Non-U.S. Gov't     Date:  2010-06-21
Journal Detail:
Title:  Infection and immunity     Volume:  78     ISSN:  1098-5522     ISO Abbreviation:  Infect. Immun.     Publication Date:  2010 Sep 
Date Detail:
Created Date:  2010-08-18     Completed Date:  2010-09-13     Revised Date:  2011-07-25    
Medline Journal Info:
Nlm Unique ID:  0246127     Medline TA:  Infect Immun     Country:  United States    
Other Details:
Languages:  eng     Pagination:  3736-43     Citation Subset:  IM    
Affiliation:
INRS-Institut Armand-Frappier, 531, Boul. des Prairies, Laval, QC, Canada H7V 1B7.
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MeSH Terms
Descriptor/Qualifier:
Active Transport, Cell Nucleus
Animals
Cell Nucleus / metabolism*
Female
Interferon Regulatory Factor-1 / genetics
Interferon-Stimulated Gene Factor 3 / analysis,  metabolism*
Interferon-gamma / physiology*
Leishmania donovani / physiology*
Macrophage Activation*
Mice
Mice, Inbred BALB C
Phosphorylation
Signal Transduction
alpha Karyopherins / metabolism*
Grant Support
ID/Acronym/Agency:
MOP-12933//Canadian Institutes of Health Research
Chemical
Reg. No./Substance:
0/Interferon Regulatory Factor-1; 0/Interferon-Stimulated Gene Factor 3; 0/KPNA1 protein, mouse; 0/alpha Karyopherins; 0/gamma interferon activation factor; 82115-62-6/Interferon-gamma
Comments/Corrections

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