Document Detail


Lateral diffusion of inositol 1,4,5-trisphosphate receptor type 1 in Purkinje cells is regulated by calcium and actin filaments.
MedLine Citation:
PMID:  20626556     Owner:  NLM     Status:  MEDLINE    
Abstract/OtherAbstract:
Inositol 1,4,5-trisphosphate receptor type 1 (IP(3) R1) is an intracellular Ca(2+) release channel that plays crucial roles in the functions of Purkinje cells. The dynamics of IP(3) R1 on the endoplasmic reticulum membrane and the distribution of IP(3) R1 in neurons are thought to be important for the spatial regulation of Ca(2+) release. In this study, we analyzed the lateral diffusion of IP(3) R1 in Purkinje cells in cerebellar slice cultures using fluorescence recovery after photobleaching. In the dendrites of Purkinje cells, IP(3) R1 showed lateral diffusion with an effective diffusion constant of approximately 0.30 μm(2) /s, and the diffusion of IP(3) R1 was negatively regulated by actin filaments. We found that actin filaments were also involved in the regulation of IP(3) R1 diffusion in the spine of Purkinje cells. Glutamate or quisqualic acid stimulation, which activates glutamate receptors and leads to a Ca(2+) transient in Purkinje cells, decreased the diffusion of IP(3) R1 and increased the density of actin in spines. These findings indicate that the neuronal activity-dependent augmentation of actin contributes to the stabilization of IP(3) R1 in spines.
Authors:
Kazumi Fukatsu; Hiroko Bannai; Takafumi Inoue; Katsuhiko Mikoshiba
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Publication Detail:
Type:  In Vitro; Journal Article; Research Support, Non-U.S. Gov't     Date:  2010-08-19
Journal Detail:
Title:  Journal of neurochemistry     Volume:  114     ISSN:  1471-4159     ISO Abbreviation:  J. Neurochem.     Publication Date:  2010 Sep 
Date Detail:
Created Date:  2010-09-24     Completed Date:  2010-10-19     Revised Date:  -    
Medline Journal Info:
Nlm Unique ID:  2985190R     Medline TA:  J Neurochem     Country:  England    
Other Details:
Languages:  eng     Pagination:  1720-33     Citation Subset:  IM    
Copyright Information:
© 2010 The Authors. Journal Compilation © 2010 International Society for Neurochemistry.
Affiliation:
Laboratory for Developmental Neurobiology, Brain Science Institute RIKEN, Wako, Saitama, Japan.
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MeSH Terms
Descriptor/Qualifier:
Actins / physiology*
Animals
Calcium / physiology*
Dendrites / drug effects,  metabolism*
Dendritic Spines / drug effects,  metabolism
Diffusion
Excitatory Amino Acid Agonists / pharmacology
Fluorescence Recovery After Photobleaching
Glutamic Acid / pharmacology
Green Fluorescent Proteins / genetics
Inositol 1,4,5-Trisphosphate Receptors / genetics,  metabolism*
Mice
Mice, Inbred ICR
Microfilaments / physiology*
Purkinje Cells / drug effects,  metabolism*,  ultrastructure
Quisqualic Acid / pharmacology
Receptors, Glutamate / physiology
Tissue Culture Techniques
Chemical
Reg. No./Substance:
0/Actins; 0/Excitatory Amino Acid Agonists; 0/Inositol 1,4,5-Trisphosphate Receptors; 0/Receptors, Glutamate; 147336-22-9/Green Fluorescent Proteins; 52809-07-1/Quisqualic Acid; 56-86-0/Glutamic Acid; 7440-70-2/Calcium

From MEDLINE®/PubMed®, a database of the U.S. National Library of Medicine


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