Document Detail


Langat flavivirus protease NS3 binds caspase-8 and induces apoptosis.
MedLine Citation:
PMID:  11991998     Owner:  NLM     Status:  MEDLINE    
Abstract/OtherAbstract:
The flavivirus NS3 protein plays an important role in the cleavage and processing of the viral polyprotein and in the synthesis of the viral RNA. NS3 recruits NS2B and NS5 proteins to form complexes possessing protease and replicase activities through protease and nucleoside triphosphatase/helicase domains. We have found that NS3 also induces apoptosis. Expression of the Langat (LGT) virus NS3 protein resulted in a cleavage of cellular DNA and reduced the viability of cells. Coexpression of NS3 with apoptotic inhibitors (CrmA and P35) and addition of caspase peptide substrates (Z-VAD-FMK and Z-IETD-FMK) to NS3-transfected cells blocked NS3-induced apoptosis. In cotransfection experiments, NS3 bound to caspase-8 and enhanced caspase-8-mediated apoptosis. NS3 and caspase-8 colocalized in the cytoplasm of transfected cells. Deletion analysis demonstrated that at least two regions of NS3 contribute to its apoptotic activities. The protease and helicase domains are each able to bind to caspase-8, while the protease domain alone induces apoptosis. The protease domain and tetrahelix region of the helicase domain are required for NS3 to augment caspase-8-mediated apoptosis. Thus, the LGT virus NS3 protein is a multifunctional protein that binds to caspase-8 and induces apoptosis.
Authors:
Grigori G Prikhod'ko; Elena A Prikhod'ko; Alexander G Pletnev; Jeffrey I Cohen
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Publication Detail:
Type:  Journal Article    
Journal Detail:
Title:  Journal of virology     Volume:  76     ISSN:  0022-538X     ISO Abbreviation:  J. Virol.     Publication Date:  2002 Jun 
Date Detail:
Created Date:  2002-05-06     Completed Date:  2002-06-10     Revised Date:  2009-11-18    
Medline Journal Info:
Nlm Unique ID:  0113724     Medline TA:  J Virol     Country:  United States    
Other Details:
Languages:  eng     Pagination:  5701-10     Citation Subset:  IM    
Affiliation:
Laboratory of Infectious Diseases, National Institute of Allergy and Infectious Diseases, National Institutes of Health, Bethesda, Maryland 20892, USA. PrikhodkoG@usa.redcross.org
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MeSH Terms
Descriptor/Qualifier:
Animals
Apoptosis*
Binding Sites
Caspase 8
Caspase 9
Caspases / genetics,  metabolism*
Cell Line, Transformed
Cercopithecus aethiops
Encephalitis Viruses, Tick-Borne / enzymology*,  genetics
Humans
Mice
RNA Helicases
Recombinant Fusion Proteins / genetics,  metabolism
Serine Endopeptidases
Tumor Cells, Cultured
Vero Cells
Viral Nonstructural Proteins / genetics,  metabolism*
Chemical
Reg. No./Substance:
0/NS3 protein, flavivirus; 0/Recombinant Fusion Proteins; 0/Viral Nonstructural Proteins; EC 2.7.7.-/RNA Helicases; EC 3.4.21.-/Serine Endopeptidases; EC 3.4.22.-/CASP8 protein, human; EC 3.4.22.-/CASP9 protein, human; EC 3.4.22.-/Casp8 protein, mouse; EC 3.4.22.-/Casp9 protein, mouse; EC 3.4.22.-/Caspase 8; EC 3.4.22.-/Caspase 9; EC 3.4.22.-/Caspases
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