Document Detail

Korean species of Aleochara Gravenhorst subgenus Xenochara Mulsant & Rey (Coleoptera, Staphylinidae, Aleocharinae).
Jump to Full Text
MedLine Citation:
PMID:  21594199     Owner:  NLM     Status:  PubMed-not-MEDLINE    
Abstract/OtherAbstract:
A taxonomic review of Aleochara Gravenhorst subgenus Xenochara Mulsant & Rey in Korea is presented. Five species are recognized, with one species, Aleochara (Baryodma) intricata Mannerheim, newly transferred to the subgenus Xenochara. Aleochara (Xenochara) asiatica Kraatz and Aleochara (Xenochara) peninsulae Bernhauer are reported for the first time in the Korean peninsula. A key, line drawings of diagnostic characters, and redescriptions of Korean Xenochara species are provided.
Authors:
Jong-Seok Park; Kee-Jeong Ahn
Related Documents :
11789869 - Successful erbium: yag laser-induced chorioretinal venous anastomosis for the managemen...
7097339 - Persisting recessus infundibuli and empty sella. case report.
21833989 - Checking reference lists to find additional studies for systematic reviews.
22263209 - Luxatio erecta: case series with review of diagnostic and management principles.
22240169 - Coherence of evidence from systematic reviews as a basis for evidence strength - a case...
22208859 - The imaging and management of nonconsecutive pars interarticularis defects: a case repo...
8677949 - Blue rubber bleb nevus syndrome.
22606429 - Postoperative bowel perforation due to heterotopic ossification (myositis ossificans tr...
11789869 - Successful erbium: yag laser-induced chorioretinal venous anastomosis for the managemen...
Publication Detail:
Type:  Journal Article     Date:  2010-10-07
Journal Detail:
Title:  ZooKeys     Volume:  -     ISSN:  1313-2970     ISO Abbreviation:  Zookeys     Publication Date:  2010  
Date Detail:
Created Date:  2011-05-19     Completed Date:  2011-07-14     Revised Date:  2013-05-29    
Medline Journal Info:
Nlm Unique ID:  101497933     Medline TA:  Zookeys     Country:  Bulgaria    
Other Details:
Languages:  eng     Pagination:  21-36     Citation Subset:  -    
Affiliation:
Department of Entomology, Louisiana State University Agricultural Center, Baton Rouge, LA, 70803, USA.
Export Citation:
APA/MLA Format     Download EndNote     Download BibTex
MeSH Terms
Descriptor/Qualifier:

From MEDLINE®/PubMed®, a database of the U.S. National Library of Medicine

Full Text
Journal Information
Journal ID (nlm-ta): Zookeys
Journal ID (publisher-id): ZooKeys
ISSN: 1313-2989
ISSN: 1313-2970
Publisher: Pensoft Publishers
Article Information
Download PDF
Jong-Seok Park, Kee-Jeong Ahn
creative-commons-attribution:
Received Day: 2 Month: 2 Year: 2010
Accepted Day: 24 Month: 3 Year: 2010
collection publication date: Year: 2010
Electronic publication date: Day: 7 Month: 10 Year: 2010
Issue: 60
First Page: 21 Last Page: 36
ID: 3088346
PubMed Id: 21594199
DOI: 10.3897/zookeys.60.404

Korean species of Aleochara Gravenhorst subgenus Xenochara Mulsant & Rey (Coleoptera, Staphylinidae, Aleocharinae)
Jong-Seok Park1
Kee-Jeong Ahn2
1Department of Entomology, Louisiana State University Agricultural Center, Baton Rouge, LA, 70803, USA
2Department of Biology, Chungnam National University, Daejeon 305-764, Republic of Korea
Correspondence: Corresponding author: Kee-Jeong Ahn (kjahn@cnu.ac.kr).
[edited-by] Academic editor: Jan Klimaszewski

The staphylinid genus [taxon-name: Aleochara] Gravenhorst includes over 400 species in 19 subgenera worldwide. Fourteen species in six subgenera are recorded in the Korean peninsula (Smetana 2004; Park and Ahn 2009). The subgenera of [taxon-name: Aleochara] have been previously diagnosed using a few characters of the antenna, mesosternal carina, pronotal pubescence and microsculpture, elytral pubescence, maxillary palpi, and genitalia (Klimaszewski 1984). However, the morphologically diverse subgenera such as [taxon-name: Xenochara] are poorly distinguished from other [taxon-name: Aleochara] subgenera by these characters (Klimaszewski 1984).

Therefore, we used characters that are more informative at the subgeneric and species levels. Our character analysis follows the methods of Sawada (1972) and Ashe (1984). The terminology for abdominal segments follows Thayer (2005).

We redescribe five Korean [taxon-name: Xenochara] species herein, and a key and line drawings of diagnostic characters of these species are also provided. The Korean specimens studied are deposited in the Chungnam National University Insect Collection (CNUIC), Daejeon, Korea.


Taxonomy
Subgenus  [taxon-name: [subgenus: Xenochara]]

[taxon-authority: Mulsant & Rey]

Type species:

[taxon-name: Aleochara decorata] Aubé.

Diagnosis.

The subgenus [taxon-name: Xenochara] can be distinguished by a combination of the following characters: body compact, robust, pubescent; antennomere 4 usually longer than wide (except [taxon-name: Aleochara tristis], transverse); carina on each side of midline of ventral surface of head present, attaining or almost attaining gular suture (arrows, Figs 1b, 2b, 3b, 4a, 5a; Klimaszewski 1984: Figs 313, 319, 321); maxillary palpomere 4 usually long (1/3 to 3/4 length of palpomere 3); labral b-seta sharpened or rounded apically (arrows, Figs 1c, 2c, 3c, 4b, 5b); mandibular internal tooth absent or weakly present; β-seta of labial palpi long (Figs 1e, 2e, 3e, 4d, 5d); mesoventrite completely or almost completely carinate (Figs 1a, 2a, 3a); pronotum evenly pubescent; spines of lateral margins of fore- and meso-tibia present but absent in meta-tibia.

Remarks.

This diagnosis is modified from Klimaszewski (1984: 35). New diagnostic characters based on mouthparts are added, and these are consistent at the subgenus level.

Key to the Aleochara (Xenochara) species from Korea
[taxon-name: [genus: Aleochara] ([subgenus: Xenochara]) [species: asiatica]]

[taxon-authority: Kraatz]

Fig. 1

Redescription.

Length 4.5–6.5 mm. Body large and robust; brownish black, antenna and legs black; elytra bicolored. Antennomeres 1–3 elongate, 4–6 longer than wide and 7–10 transverse. Mouthparts. Labrum transverse, bearing approximately 11 small, setae, and approximately 21 long setae, a-seta, b-seta, and pores present; b-seta rounded apically (arrow indicates b-seta, Fig. 1c). Labium with pseudopores in median area, approximately 2 real pores and pseudopores present in lateral area; a pair of basal pores present (Fig. 1d). Ligula with approximately 4 pairs of small setae apically (Fig. 1d). Labial palpi with large a-, b-, and f-seta of 12 setae present (a–h, α–δ); long β-seta present in the middle of palpomere 1; d-seta higher than c-seta (Fig. 1e). Mentum transverse, bearing 4 pairs of main setae (b, u, v, w), and 6 extra setae, and pores present (Fig. 1f). Thorax. Mesoventrite completely carinate (arrow, Fig. 1a). Elytra with round latero-posterior margin. Abdomen. Male and female abdominal tergite VIII with many short setae and pores; posterior margin deeply emarginate and undulate on male (Fig. 1g, h). Male and female abdominal sternite VIII with many short setae and pores, margin rounded and female with many small setae on posterior margin (Fig. 1i, j). Genitalia. Median lobe as in Figs 1l and m. Paramere without fovea in hinge zone (Fig. 1k). Spermatheca with duct coiled (Fig. 1n).

Material examined.

1♂, Daeheungsa-temple, Samsan-myeon, Haenam-gun, Jeonnam Prov., Korea, 23.IV.1983, Y. B. Cho; 4♂♂ 2♀♀, Jeju Prov., Korea, 22.VII.1985, Y.B. Cho (1♂ 1♀, on slide); 1♀, Donnaeko, Sanghyo-dong, Seogwipo-city, Jeju Prov., Korea, 18.X.1985, K.-S. Lee; 1♀, Donnaeko, Sanghyo-dong, Seogwipo-city, Jeju Prov., Korea, 22.X.1985, K.-S. Lee; 1♀, Jeongbangpolpo-waterfall, Donghong-dong, Seogwipo-city, Jeju Prov., Korea, 29.X.1985, K.-S. Lee; 1♂, Gasi-ri, Pyoseon-myeon, Namjeju-gun, Jeju Prov., Korea, 5.V.1985, K.-S. Lee; 2, Japan, G. Lewis, 1910–320, Nagasaki, 22.V–3.VI.81; 2, Japan, G. Lewis, 1910–320, Kumamoto, 23.IV–26.IV.81; 3, Japan, G. Lewis, 1905–313 (1, Holotype of [taxon-name: Aleochara japonica] Sharp, deposited in the Natural History Museum, London); 1, Japan, G. Lewis, 1910–320.

Distribution.

China, India, Japan, Korea, Nepal, Taiwan (see Smetana, 2004: 356).

Remarks.

This species is a new record for the Korean peninsula.

[taxon-name: [genus: Aleochara] ([subgenus: Xenochara]) [species: intricata]]

[taxon-authority: Mannerheim]

Fig. 2

Redescription.

Length 3.5–6.0 mm. Body large and robust; brownish black; antennomeres 1–3 and legs brown; elytra yellow to yellowish brown and bicolored. Antennomeres 1–3 elongate, 4 longer than wide, 5–8 weakly transverse and 9–10 transverse. Mouthparts. Labrum transverse, bearing approximately 8 small, setae, and approximately 26 long setae, a-seta, b-seta, and pores present; b-seta rounded apically (arrow indicates b-seta, Fig. 2c). Labium with pseudopores in median area; approximately 3 real pores and pseudopores present in lateral area; pair of basal pores present (Fig. 2d). Ligula with approximately 4 pairs of small setae apically (Fig. 2d). Labial palpi with large a-, b-, and f-seta of 12 setae (a–h, α–δ) present; β-seta close to twin pores (tp); d- and c-seta at same level (Fig. 2e). Mentum transverse, bearing 4 pairs of main setae (b, u, v, w), and 4 extra setae, and pores present (Fig. 2f). Thorax. Mesoventrite completely carinate (arrow, Fig. 2a). Elytra with round latero-posterior margin. Abdomen. Abdominal tergite VIII with many short setae and pores; posterior margin weakly emarginate (Fig. 2g). Abdominal sternite VIII with many short setae and pores; posterior margin rounded (Fig. 2h, i). Genitalia. Median lobe as in Figs 2k and l. Paramere without fovea in hinge zone (Fig. 2j). Spermatheca as in Fig. 2m.

Material examined.

7, Hol-ri, Ganseong-eup, Goseong-gun, Gangwon Prov., Korea, 31.VIII–1.IX.1984, Y.-S. Kim (2♂♂2♀♀, on slide); 1, Gohan-ri, Sabuk-eup, Gangwon Prov., Korea, 27.IV.1985, J.-I. Kim; 1, Deakwan-ryeong, Gangwon Prov., Korea, 28.VI.1984, Y.-S. Kim; 2, Oigapyeong, Inje-gun, Gangwon Prov., Korea, 26.V.1986, Y.-S. Kim; 1, Baekdamsa-temple, Inje-gun, Gangwon Prov., Korea, 26.V.1986, Y.-S. Kim; 1, Dammaeul, Cheongsong, Gyeongbuk Prov., Korea, 24.VI.1988, K.-S. Jang; 1, Tonghan, Anjeong, Jeonbuk Prov., Korea, 5.VI.1988, Y.-S. Kim; 6, Seilles, carriere 1, 11/19 VIII 1945, G. Fagel; 2, Anseremme car, vers Freyr, 26 VIII 1946, G. Fagel; 2, Abruzzo, A. colomba, Italy, 26.VII.1894, P Fiori; 12, Lazio, Roma, Lirezzi, Italy, A. Fiori.

Distribution.

Korea, Asia, Europe, North Africa (see Smetana, 2004: 354).

Remarks.

This species is a new record for South Korea.

[taxon-name: [genus: Aleochara] ([subgenus: Xenochara]) [species: peninsulae]]

[taxon-authority: Bernhauer]

Fig. 3

Holotype examined.

Male mounted on card, with mouthparts, aedeagus and abdominal apex (segment VIII+) mounted in balsam on two transparent cards, labeled as follows: “Shimabara Unzen 2200F 2. 8. 34. Suenson” [printed]; “peninsulae Brnh. Typus un. [taxon-name: Polychara]” [handwritten yellow label]; “peninsulae Brnh. Typus unic.” [handwritten white label]; “Chicago NHMus M. Bernhauer Collection” [printed]; “HOLOTYPE [taxon-name: Aleochara peninsulae] Bernhauer, 1936 teste Park & Ahn 2007” . Deposited in the Field Museum of Natural History (FMNH), Chicago, USA.

Redescription.

Length 3.8–5.4 mm. Body compact and robust; reddish black; antenna, elytra, and legs brownish black; elytra bicolored. Antennomeres 1–3 elongate, 4 longer than wide, 5 subquadrate, and 6–10 transverse. Mouthparts. Labrum transverse, bearing approximately 8 small, setae, and approximately 19 long setae, a-seta, b-seta and pores present; b-seta acute (arrow indicates b-seta, Fig. 3c). Labium with pseudopores in median area; approximately 3 pores and pseudopores present in lateral area; a pair of distal setae present (arrow indicates distal seta, Fig. 3d). Ligula with approximately 3 pairs of small setae apically (Fig. 3d). Labial palpi with large a-, b-, and f-seta of 12 setae present (a–h, α–δ); β-seta close to twin pores (tp); d-seta higher than c-seta (Fig. 3e). Mentum transverse, bearing 4 pairs of main setae (b, u, v, w), and approximately 2 extra setae, and pores present (Fig. 3f). Thorax. Mesoventrite completely carinate (arrow, Fig. 3a). Elytral latero-posterior margin emarginate. Abdomen. Abdominal tergite VIII with many short setae and pores; posterior margin emarginate; undulate on male (Fig. 3g, h). Male abdominal sternite VIII with many short setae and pores; posterior margin produced on male (arrow, Fig. 3i) and rounded on female (Fig. 3j). Genitalia. Median lobe as in Figs 3l and m. Paramere with fovea in hinge zone (arrow, Fig. 3k). Spermatheca as in Fig. 3n.

Material examined.

2, Gyeongsang Univ., Gajoa-Dong, Jinju-city, Gyeongnam Prov., Korea, 26.III.2003, C.-S. Lim, ex bait trap; 2, Gyeongsang Univ., Gajoa-Dong, Jinju-city, Gyeongnam Prov., Korea, 2 VI 2003, C.-S. Lim, ex bait trap; 1, Gyeongsang Univ., Gajoa-Dong, Jinju-city, Gyeongnam Prov., Korea, 27.III.2003, C.-S. Lim, ex bait trap; 2, Gyeongsang Univ., Gajoa-Dong, Jinju-city, Gyeongnam Prov., Korea, 22.III.2003, C.-S. Lim, ex bait trap; 1, Gyeongsang Univ., Gajoa-Dong, Jinju-city, Gyeongnam Prov., Korea, 12.III.2003, C.-S. Lim, ex bait trap; 1, Gyeongsang Univ., Gajoa-Dong, Jinju-city, Gyeongnam Prov., Korea, 7.IV.2003, C.-S. Lim, ex bait trap; 1, Gyeongsang Univ., Gajoa-Dong, Jinju-city, Gyeongnam Prov., Korea, 15.III.2003, C.-S. Lim, ex bait trap; 1, Gyeongsang Univ., Gajoa-Dong, Jinju-city, Gyeongnam Prov., Korea, 24.III.2003, C.-S. Lim, ex bait trap; 1, Gyeongsang Univ., Gajoa-Dong, Jinju-city, Gyeongnam Prov., Korea, 29.V.2003, C.-S. Lim, ex bait trap; 1, Gyeongsang Univ., Gajoa-Dong, Jinju-city, Gyeongnam Prov., Korea, 30.III.2003, C.-S. Lim, ex bait trap; 1, near 1100m Rest Area, Jeju Prov., Korea, 30.V–17.VI.2003, Y.-B. Cho, ex bait trap.

Distribution.

Japan, Korea.

Remarks.

This species is a new record for the Korean peninsula.

[taxon-name: [genus: Aleochara] ([subgenus: Xenochara]) [species: puberula]]

[taxon-authority: Klug]

Fig. 4

Redescription

Length 3.5–5.5 mm. Body reddish brown; antennomeres 1–3, elytra, and legs brown; elytra bicolored. Antennomeres 1–3 elongate, 4 longer than wide, 5 subquadrate, and 6–10 transverse. Mouthparts. Labrum transverse, bearing approximately 11 small, setae, and approximately 19 long setae, a-seta, b-seta, and pores present; b-set a rounded at tip (arrow indicates b-seta, Fig. 4b). Labium with pseudopores in median area; approximately 3 real pores and pseudopores present in lateral area; a pair of basal pores present (Fig. 4c). Ligula with approximately 2 pairs of small setae apically (Fig. 4c). Labial palpi with large a-, b-, and f-seta of 12 setae present (a–h, α–δ); β-seta close to twin pores (tp); c-seta higher than d-seta (Fig. 4d). Mentum transverse, bearing 4 pairs of main setae (b, u, v, w), and 4 extra setae, and pores present (Fig. 4e). Thorax. Elytral latero-posterior margin emarginate. Abdomen. Abdominal tergite VIII with many short setae and pores; posterior margin weakly emarginate (Fig. 4f). Abdominal sternite VIII with many short setae and pores; apical margin rounded on male (Fig. 4g) and truncated on female (Fig. 4h). Genitalia. Median lobe as in Figs 4j and k. Paramere with fovea absent in hinge zone (Fig. 4i). Spermatheca coiled in duct (Fig. 4l).

Material examined.

2, Seoguipo-city, Jeju Prov., Korea, 18.VI.1985, K.-S. Lee; 1, Anseong, Muju-gun, Jeonbuk Prov., Korea, 5.VI.1988, G.-S. Jang; 22, Reunion 22–23.I.1992, Ravine de St. Gilles Bassin Cormoran, J. Janaj lgt; 1, Philippinen, Manila, Luy, 2.XI.1914; 1, Argentina, Prov. Tucuman, 450m, I 1905, Steinbach; 1, N. Palawan, Bakuit, 12.XI–22.XII 1913, Bottcher; 4, Sud algérien: Mrhaier 120 Km S de Biskra, 14.V.1954, G. Fagel; 4, China, B. M. 1980–491, P. M. Hammond, Guangdong, Guangzhou, Baiyunshan, 27.IX.80; 4, China, B. M. 1980–491, P. M. Hammond, Guizhou, 20m, S. Guilin, 22.IX.80; 1, Japan, Honshu, B. M. 1980–492, P. M. Hammond, Nara, ft of Mt. Kasuga, 20.VIII.80; 2, Japan, Sharp Coll., 1905–313; 4, Japan, G. Lewis, 1910–320.

Distribution.

Japan, Korea, Asia, Europe, North Africa, North America (see Smetana, 2004: 360).

[taxon-name: [genus: Aleochara] ([subgenus: Xenochara]) [species: tristis]]

[taxon-authority: Gravenhorst]

Fig. 5

Redescription.

Length 3.7–6.4 mm. Body black; antenna and legs reddish black; elytra yellow to yellowish brown and bicolored. Antennomeres 1–3 elongate, and 4–10 transverse. Mouthparts. Labrum transverse, bearing approximately 9 small, setae, and approximately 17 long setae, a-seta, b-seta, and pores resent; b-seta acute (arrow indicates b-seta, Fig. 5b). Labium with pseudopores in median area; about 3 real pores and pseudopores present in lateral area; a pair of distal setae present (arrow indicates distal seta, Fig. 5c). Ligula with approximately 3 pairs of small setae apically (Fig. 5c). Labial palpi with large a-, b-, and f-seta of 12 setae present (a–h, α–δ); β-seta close to twin pores (tp); c- and d-seta same level (Fig. 5d). Mentum transverse, bearing 4 pairs of main setae (b, u, v, w), and 13 extra setae, and pores present (Fig. 5e). Thorax. Elytra with round latero-posterior margin. Abdomen. Abdominal tergite VIII with many short setae and pores; posterior margin emarginate (Fig. 5f). Abdominal sternite VIII with many short setae and pores; apical margin rounded (Fig. 5g, h). Genitalia. Median lobe with coiled flagellum (arrow, Figs 5j and k). Paramere with fovea in hinge zone (arrow, Fig. 5i). Spermathecal duct coiled (Fig. 5l).

Material examined

1, Russia, Tadshikistan, 1981, Duechanbe env, on light, 16–18.VI.1981, K. Majer; 36, Russia, Tadshikistan, Umg. Duechanbe env, 800–1200m, 4–14.IX.1983, U. Arnold.

Distribution.

Korea, Asia, Europe, North Africa, North America (see Smetana, 2004: 361).

Remarks.

See Klimaszewski (1984: 39) for reason why the older names [taxon-name: Staphylinus bipunctata] Olivier or [taxon-name: Staphylinus geometrica] Schrank do not have priority. We used Russian specimens for the redescription.


Acknowledgements

We thank Mr. James H. Boone and Dr. Alfred F. Newton (FMNH, Chicago) for the loan of type specimen and encouragements. We also thank Chris Carlton, Matthew Gimmel (Louisiana State Univ., Baton Rouge), and anonymous reviewers for helpful comments. This publication was approved by the Director, Louisiana Agricultural Experiment Station as manuscript number 2010-234-4180. This research was supported by the project on survey and excavation of Korean indigenous species of the National Institute of Biological Resources (NIBR) under the Ministry of Environment, Korea.


References
Ashe JS. (Year: 1984) Generic revision of the subtribe Gyrophaenini (Coleoptera: Staphylinidae: Aleocharinae) with a review of the described subgenera and major features of evolution.Quaestiones Entomologicae20:129-349
Assing V. (Year: 2007a) New species and additional records of Staphylinidae from Turkey V (Coleoptera).Stuttgarter Beiträge zur Naturkunde, Serie A (Biologie)700:1-64
Assing V. (Year: 2007b) On the Aleocharini of Turkey, with notes on some species from adjacent regions (Coleoptera: Staphylinidae, Aleocharinae).Beiträge zur Entomologie57 (1):177-209
Aubé C. (Year: 1850) Description de quelques insectes coléoptères appartenant à l'Europe et à l'Algérie.Annales de la Société Entomologique de France (2)8:299-346
Bernhauer M. (Year: 1901) Die Staphyliniden der paläarktischen Fauna.Verhandlungen der Kaiserlich-Königlichen Zoologisch-Botanischen Gesellschaft in Wien51:430-506
Bernhauer M. (Year: 1936) Neuheiten der palaearktischen Staphyliniden fauna.Pubblicazioni del Museo Entomologico «Pietro Rossi» Duino1:237-325
Bernhauer M,Scheerpeltz O. (Year: 1926) Staphylinidae VI. (Pars 82). In: Junk W, Schenkling S (Eds) Coleopterorum Catalogus. Volumen 5. Staphylinidae.W Junk, Berlin, 988 pp.
Cameron M. (Year: 1939) Fauna of British India including Ceylon and Burma. Coleoptera Staphylinidae. Volume IVParts I & II. Taylor and Francis, London, xviii + 691 pp.
Casey TL. (Year: 1894) Coleopterological notices. V.Annals of the New York Academy of Sciences7 (“1893”): 281–606
Eichelbaum F. (Year: 1912) Die von Herrn Dr. Christoph Schröder in den Parehbergen und auf dem Kilimandjaro im Januar 1906 gesammelten Staphylinidae (darunter zwei neue Philonthusarten).Berliner Entomologische Zeitschrift56 (“1911”): 173–176
Erichson WF. (Year: 1839) Genera et species Staphylinorum insectorum coleopterorum familiae 1.F.H. Morin, Berlin, 954 pp.
Fauvel A. (Year: 1863) In: Chevrolat A,Fauvel A. Coléoptères de l'Ile de Cuba. Notes, synonymiques et descriptions d'espèces nouvelles. Annales de la Société Entomologique de France (4)3: 427–446
Fenyes A. (Year: 1920) Coleoptera Fam. Staphylinidae subfam. Aleocharinae. Genera Insectorum.173: 111–414
Fowler WW. (Year: 1888) The Coleoptera of the British Islands. A descriptive account of the families, genera, and species indigenous to Great Britain and Ireland, notes as to localities, habtiats, etc. Volume II, L Reeve & Co., London, 444 pp.
Ganglbauer L. (Year: 1895) Die Käfer von Mitteleuropa. Die Käfer der österreichisch-ungarischen Monarchie, Deutschland, der Schweiz, sowie des französischen und italienischen Alpengebietes. Zweiter Band. Famielienreihe Staphylinoidea. 1. Theil: Staphylinidae, Pselaphidae, Carl Gerold's Sohn, Wien, 881 pp.
Gravenhorst JKC. (Year: 1806) Monographia Coleopterorum Micropterorum, Henricus Dieterich, Göttingen, 236 pp.
Heer O. (Year: 1839) Fascicle II. In: Fauna Coleopterorum Helvetica. Pars I. Orelii, Fuesslini & Soc., Zürich, 652 pp.
Jekel H. (Year: 1873) Coleoptera Jekeliana, adjecta Eleutheratorum Bibliotheca. Énumération systematique et synonymie des coléoptères européens et exotiques composant la collection de Henri Jekel. Observations critiques. Descriptions d’espèces nouvelles. Reproductions et traduction de genres et espèces publiés dans des ouvrages francais et étrangers de manière à former insensiblement la Bibliothèque ou Coléoptèriste. Livraison I, E. Deyrolle Fils, Paris, 96 pp.
Klimaszewski J. (Year: 1984) A Revision of the genus Aleochara Gravenhorst of America North of Mexico (Coleoptera: Staphylinidae: Aleocharinae).Memoirs of the Entomological Society of Canada129:1-211
Klug JCF. (Year: 1832) Bericht über eine auf Madagascar veranstaltete Sammlung von Insecten aus der Ordnung Coleoptera.Abhandlungen der Königlichen Akademie der Wissenschaften zu Berlin1832:91-223
Kraatz G. (Year: 1859) Die Staphylinen-Fauna von Ostindien, insbesondere der Insel Ceylan.Archiv für Naturgeschichte25:1-196
Lohse GA. (Year: 1974) 23. Familie: Staphylinidae. In: Freude H,Harde KW,Lohse GA. (Eds) Die Käfer Mitteleuropas. Band 5, Staphylinidae II (Hypocyphtinae und Aleocharinae). Pselaphidae.Goecke & Evers, Krefeld, 304 pp.
MacLeay WJ. (Year: 1873) Notes on a collection of insects from Gayndah.The Transactions of the Entomological Society of New South Wales2:79-205
Mannerheim CG. (Year: 1830) Précis d'un nouvel arrangement de la famille des brachélytres de l'ordre des insectes coléoptères, St. Petersburg, 87 pp.
Ménétriés E. (Year: 1832) Catalogue raisonné des objets de zoologie recueillis dans un voyage au Caucase et jusqu'aux frontières actuelles de la Perse entrepris par ordre de S. M. l'Empereur, L'Académie Impériale des Sciences, St. Petersburg, 271 pp.
Miller L. (Year: 1853) Drei neue Staphylinen der Wiener Gegend. Verhandlungen der Zoologisch-Botanischen Vereins in Wien 2 (“1852”): 26–28
Motschulsky V. (Year: 1858) Énumeration des nouvelles espèces des de coléoptères rapportés de ses voyages.Bulletin de la Société Impériale des Naturalistes de Moscou31 (3):204-264
Mulsant E,Rey C. (Year: 1874) Histoire naturelle des coléoptères de France. Brévipennes. Aléochariens. (Suite). - Aléocharaires, Deyrolle, Paris, 565 pp.
Olivier AG. (Year: 1795) Entomologie, ou histoire naturelle des insectes, avec leur caractères génériques et spécifiques, leur description, leur synonymie, et leur figure illuminée. Coléoptères, Lanneau, Paris, 557 pp.
Pace R. (Year: 2001) Aleocharinae nuove o poco note dell’India (Coleoptera, Staphylinidae) (165° Contributo alla conoscenza delle Aleocharinae).Nouvelle Revue d’Entomologie18 (1):31-47
Palm T. (Year: 1972) Skalbaggar. Coleoptera. Kortvingar: Fam. Staphylinidae UnderFam. Aleocharinae (Aleuonota-Tinotus).Svensk Insektfauna53:310-467
Park JS,Ahn KJ. (Year: 2009) Synonymy of Aleochara (Aleochara) claviger and A. niponensis (Coleoptera: Staphylinidae: Aleocharinae), and first record of A. claviger in Korea.Florida Entomologist92 (3):521-523
Penecke KA. (Year: 1901) Coleopterologische Miscellen. II.Wiener Entomologische Zeitung20:11-21
Portevin G. (Year: 1929) Histoire naturelle des Coléoptères de France. Tome I. Adephaga. - Polyphaga: Staphylinoidea.Encyclopédie entomologique. Série A. Tome XII, Paul Lechevalier, Paris, 649 pp.
Sawada K. (Year: 1972) Methodological research in the taxonomy of Aleocharinae.Contributions from the Biological Laboratory Kyoto University24 (1):31-59
Schrank FP. (Year: 1798) Fauna Boica. Durchgedachte Geschichte der in Bairen einhemischen und zahmen Thiere, Vo. 1Stein, Nürnberg, 720 pp.
Seevers CH. (Year: 1978) A generic and tribal revision of the North American Aleocharinae (Coleoptera: Staphylinidae).Fieldiana Zoology71:1-289
Sharp DS. (Year: 1874) The Staphylinidae of Japan.The Transactions of the Entomological Society of London1874:1-103
Smetana A. (Year: 2004) Aleocharinae. In: Löbl I,Smetana A. (Eds) Catalogue of Palaearctic Coleoptera Volume 2, Hydrophiloidea, Histeroidea, Staphylinoidea.Apollo Books, Stenstrup, 353–295 pp.
Stephens JF. (Year: 1832) Illustrations of British entomology; or, a synopsis of indigenous insects, containing their generic and specific distinctions; with account of their metamorphoses, times of appearance, times of appearance, localities, food, and economy, as far as practicable. Mandibulata. Vol. VBaldwin & Cradock, London, 448 pp + pls. 24–27.
Thayer MK. (Year: 2005) Staphylinidae Latreille, 1802. In: Beutel RG,Leschen AB. (Eds) Coleoptera, beetles volume 1: morphology and systematics (Archostemata, Adephaga, Myxophaga, Polyphaga partim).Walter de Gruyter, Berlin, New York, 296–344
Welch RC. (Year: 1997) The British species of the genus Aleochara Gravenhorst (Staphylinidae).Coleopterist6 (1):1-45
Wollaston TV. (Year: 1854) Insecta Maderensia; being an account of the insects of the islands of the Madeiran group.V.J. Voorst, London, 634 pp.

Figures

[Figure ID: F1]
Figure 1. 

[taxon-name: Aleochara (Xenochara) asiatica]. a meso- and metaventrite, ventral aspect b head, ventral aspect c labrum, dorsal aspect d labium, ventral aspect e labial palpus, ventral aspect f mentum, ventral aspect g male tergite VIII, dorsal aspect h female tergite VIII, dorsal aspect i male sternite VIII, ventral aspect j female sternite VIII, ventral aspect k paramere, lateral aspect l median lobe, lateral aspect m median lobe, dorsal aspect n spermatheca. Scale bars = 0.1 mm.



[Figure ID: F2]
Figure 2. 

[taxon-name: Aleochara (Xenochara) intricata]. a meso- and metaventrite, ventral aspect b head, ventral aspect c labrum, dorsal aspect d labium, ventral aspect e labial palpus, ventral aspect f mentum, ventral aspect g tergite VIII, dorsal aspect h male sternite VIII, ventral aspect i female sternite VIII, ventral aspect j paramere, lateral aspect k median lobe, lateral aspect l median lobe, dorsal aspect m spermatheca. Scale bars = 0.1 mm.



[Figure ID: F3]
Figure 3. 

[taxon-name: Aleochara (Xenochara) peninsulae]. a meso- and metaventrite, ventral aspect b head, ventral aspect c labrum, dorsal aspect d labium, ventral aspect e labial palpus, ventral aspect f mentum, ventral aspect g male tergite VIII, dorsal aspect h female tergite VIII, dorsal aspect i male sternite VIII, ventral aspect j female sternite VIII, ventral aspect k paramere, lateral aspect l median lobe, lateral aspect m median lobe, dorsal aspect n spermatheca. Scale bars = 0.1 mm.



[Figure ID: F4]
Figure 4. 

[taxon-name: Aleochara (Xenochara) puberula]. a head, ventral aspect b labrum, dorsal aspect c labium, ventral aspect d labial palpus, ventral aspect e mentum, ventral aspect f tergite VIII, dorsal aspect g male sternite VIII, ventral aspect h female sternite VIII, ventral aspect i paramere, lateral aspect j median lobe, lateral aspect k median lobe, dorsal aspect l spermatheca. Scale bars = 0.1 mm.



[Figure ID: F5]
Figure 5. 

[taxon-name: Aleochara (Xenochara) tristis]. a head, ventral aspect b labrum, dorsal aspect c labium, ventral aspect d labial palpus, ventral aspect e mentum, ventral aspect f tergite VIII, dorsal aspect g male sternite VIII, ventral aspect h female sternite VIII, ventral aspect i paramere, lateral aspect j median lobe, lateral aspect k median lobe, dorsal aspect l spermatheca. Scale bars = 0.1 mm.



Tables
[TableWrap ID: d33e349]
1. Elytra with emarginate latero-posterior margins 2
Elytra with rounded latero-posterior margins 3
2. Labral b-seta acute (arrow, Fig. 3c), labium with a pair of distal setae (arrow, Fig. 3d), male abdominal tergite VIII with undulate posterior margin (secondary sexual dimorphism; Fig. 3g), male abdominal sternite VIII with produced posterior margin (secondary sexual dimorphism; arrow, Fig. 3i), paramere with fovea in hinge zone (arrow, Fig. 3k), spermathecal duct not coiled (Fig. 3n) [taxon-name: Aleochara peninsulae] Bernhauer
Labral b-seta rounded at tip (arrow, Fig. 4b), labium without distal setae (Fig. 4c), male abdominal tergite VIII without undulate posterior margin (Fig. 4f), male abdominal sternite VIII with rounded posterior margin (Fig. 4g), paramere without fovea in hinge zone (Fig. 4j), spermathecal duct coiled (Fig. 4l) [taxon-name: Aleochara puberula] Klug
3. Antennomeres 5–6 longer than wide, abdominal tergite VIII with deeply emarginate posterior margin (Figs 1g, 1h), male tergite VIII with undulate posterior margin (secondary sexual dimorphism; Fig. 1g) [taxon-name: Aleochara asiatica] Kraatz
Antennomeres 5–6 transverse, abdominal tergite VIII with weakly emarginate posterior margin (Figs 2g, 5f), male tergite VIII without undulate posterior margin (Figs 2g, 5f) 4
4. Antennomere 4 longer than wide, labral b-seta rounded at tip (arrow, Fig. 2c), labium without distal setae (Fig. 2d), paramere without fovea in hinge zone (Fig. 2j), median lobe without coiled flagellum (Fig. 2k), spermathecal duct not coiled (Fig. 2m) [taxon-name: Aleochara intricata] Mannerheim
Antennomere 4 transverse, labral b-seta acute (arrow, Fig. 5b), labium with a pair of distal setae (arrow, Fig. 5c), paramere with fovea in hinge zone (arrow, Fig. 5i), median lobe with coiled flagellum (arrow, Fig. 5j), spermathecal duct coiled (Fig. 5l) [taxon-name: Aleochara tristis] Gravenhorst


Article Categories:
  • Article

Keywords: Keywords  Aleocharini, Aleochara, Xenochara, redescription, Korea.

Previous Document:  Parasabella Bush, 1905, replacement name for the polychaete genus Demonax Kinberg, 1867 (Annelida, P...
Next Document:  A review of the non-metallic Osmia (Melanosmia) found in North America, with additional notes on pal...