Document Detail


Interferon regulatory factor-8-driven myeloid differentiation is regulated by 12/15-lipoxygenase-mediated redox signaling.
MedLine Citation:
PMID:  20647030     Owner:  NLM     Status:  MEDLINE    
Abstract/OtherAbstract:
OBJECTIVE: Several transcription factors determine the cell fate decision between granulocytes and monocytes, but the upstream signal transduction pathways that govern myelopoiesis are largely unknown. Based on our observation of aberrant myeloid cell representation in hematopoietic tissues of 12/15-lipoxygenase (12/15-LOX)-deficient (Alox15) mice, we tested the hypothesis that polyunsaturated fatty acid metabolism regulates myelopoiesis.
MATERIALS AND METHODS: Multicolor flow cytometric analysis and methylcellulose assays were used to compare myelopoiesis and the differentiative capacity of progenitors from Alox15 and wild-type mice. Furthermore, we elucidated the mechanism by which 12/15-LOX is involved in regulation of myelopoiesis.
RESULTS: Granulopoiesis in Alox15 mice is increased while monopoiesis is reduced. Moreover, there is an accumulation of granulocyte-macrophage progenitors that exhibit defective differentiation. Mechanistically, we demonstrate that transcriptional activity of interferon regulatory factor-8 (Irf8), which regulates myelopoiesis, is impaired in Alox15 progenitors and bone marrow-derived macrophages due to loss of 12/15-LOX-mediated redox regulation of Irf8 nuclear accumulation. Restoration of redox signaling in Alox15 bone marrow cells and granulocyte-macrophage progenitors reversed the defect in myeloid differentiation.
CONCLUSIONS: These data establish 12/15-LOX-mediated redox signaling as a novel regulator of myelopoiesis and Irf8.
Authors:
Michelle Kinder; James E Thompson; Cong Wei; Suresh G Shelat; Ian A Blair; Martin Carroll; Ellen Puré
Publication Detail:
Type:  Journal Article; Research Support, N.I.H., Extramural; Research Support, Non-U.S. Gov't     Date:  2010-07-18
Journal Detail:
Title:  Experimental hematology     Volume:  38     ISSN:  1873-2399     ISO Abbreviation:  Exp. Hematol.     Publication Date:  2010 Nov 
Date Detail:
Created Date:  2010-10-22     Completed Date:  2010-12-30     Revised Date:  2011-06-14    
Medline Journal Info:
Nlm Unique ID:  0402313     Medline TA:  Exp Hematol     Country:  Netherlands    
Other Details:
Languages:  eng     Pagination:  1036-1046.e1-4     Citation Subset:  IM    
Copyright Information:
Copyright © 2010 ISEH - Society for Hematology and Stem Cells. Published by Elsevier Inc. All rights reserved.
Affiliation:
The Wistar Institute, Philadelphia, PA 19104, USA.
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MeSH Terms
Descriptor/Qualifier:
Animals
Arachidonate 12-Lipoxygenase / deficiency*,  genetics
Arachidonate 15-Lipoxygenase / deficiency*,  genetics
Cell Differentiation / drug effects
Cell Nucleus / metabolism
Erythropoietin / pharmacology
Female
Flow Cytometry
Granulocyte-Macrophage Colony-Stimulating Factor / pharmacology
Granulocyte-Macrophage Progenitor Cells / cytology,  metabolism
Granulocytes / cytology,  metabolism
Immunoblotting
Interferon Regulatory Factors / metabolism*
Interleukin-3 / pharmacology
Interleukin-6 / pharmacology
Macrophages / cytology,  metabolism
Male
Mice
Mice, Inbred C57BL
Mice, Knockout
Monocytes / cytology,  metabolism
Myeloid Progenitor Cells / cytology,  metabolism*
Myelopoiesis / drug effects
Oxidation-Reduction
Reactive Oxygen Species / metabolism
Signal Transduction*
Grant Support
ID/Acronym/Agency:
P30 ES013508-039001/ES/NIEHS NIH HHS; P30CA10815/CA/NCI NIH HHS; P30ES013508/ES/NIEHS NIH HHS; R01 CA091016-09/CA/NCI NIH HHS; R01CA091016/CA/NCI NIH HHS; T32CA09171/CA/NCI NIH HHS; T32GM07229/GM/NIGMS NIH HHS
Chemical
Reg. No./Substance:
0/12-15-lipoxygenase; 0/Interferon Regulatory Factors; 0/Interleukin-3; 0/Interleukin-6; 0/Reactive Oxygen Species; 0/interferon regulatory factor-8; 11096-26-7/Erythropoietin; 83869-56-1/Granulocyte-Macrophage Colony-Stimulating Factor; EC 1.13.11.31/Arachidonate 12-Lipoxygenase; EC 1.13.11.33/Arachidonate 15-Lipoxygenase

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