Document Detail


Involvement of matrix metalloproteinases in the adipose conversion of 3T3-L1 preadipocytes.
MedLine Citation:
PMID:  12049638     Owner:  NLM     Status:  MEDLINE    
Abstract/OtherAbstract:
When mouse 3T3-L1 preadipocytes are induced to differentiate into adipocytes, they change from an extended fibroblast-like morphology to a rounded one. This change most likely occurs through extracellular matrix remodelling, a process known to be mediated in part by matrix metalloproteinases (MMPs). In this study, we have shown by semi-quantitative reverse transcriptase-PCR, zymographic and immunoblot analysis that MMP-2, MMP-9 and membrane type 1 (MT1)-MMP are regulated during adipose conversion. To assess the importance of MMPs for adipocytic differentiation we have used MMP-specific inhibitors as well as neutralizing antibodies. Treatment of 3T3-L1 preadipocytes with the broad MMP inhibitor Ilomastat or the more restricted MMP-2 Inhibitor I prevented their differentiation into adipocytes in a dose-dependent manner, as evidenced by absence of triglyceride accumulation. Inhibitor treatment prevented the fibronectin-network degradation, as well as the induction of the genes for peroxisome-proliferator-activated receptor gamma and adipsin, two adipocyte phenotype markers. Inhibitor treatment was effective when applied during the early stages of adipocytic conversion, whereas inhibitor treatment during later stages had little effect. Inhibitor treatment did not inhibit clonal mitotic expansion; nor did it affect the expression pattern of the adipogenic transcription factor CCAAT/enhancer-binding protein beta (C/EBPbeta) or its nuclear translocation. It did, however, markedly reduce C/EBPbeta DNA-binding capacity. Taken together, these results suggest that MMPs, and notably MMP-2 and MMP-9, may be necessary mediators of adipocytic differentiation of 3T3-L1 cells.
Authors:
Gilles Croissandeau; Michel Chrétien; Majambu Mbikay
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Publication Detail:
Type:  Journal Article; Research Support, Non-U.S. Gov't    
Journal Detail:
Title:  The Biochemical journal     Volume:  364     ISSN:  0264-6021     ISO Abbreviation:  Biochem. J.     Publication Date:  2002 Jun 
Date Detail:
Created Date:  2002-06-06     Completed Date:  2002-08-15     Revised Date:  2009-11-18    
Medline Journal Info:
Nlm Unique ID:  2984726R     Medline TA:  Biochem J     Country:  England    
Other Details:
Languages:  eng     Pagination:  739-46     Citation Subset:  IM    
Affiliation:
Diseases of Aging Program, Ottawa Health Research Institute, University of Ottawa, Ottawa, Ontario K1Y 4K9, Canada.
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MeSH Terms
Descriptor/Qualifier:
3T3 Cells
Adipocytes / cytology*,  enzymology
Animals
Antibodies / pharmacology
CCAAT-Enhancer-Binding Protein-beta / metabolism
Cell Differentiation / drug effects
Indoles / pharmacology
Kinetics
Matrix Metalloproteinase 14
Matrix Metalloproteinase 2 / metabolism
Matrix Metalloproteinase 9 / metabolism
Matrix Metalloproteinases / metabolism*
Matrix Metalloproteinases, Membrane-Associated
Metalloendopeptidases / metabolism
Mice
Protease Inhibitors / pharmacology
Reverse Transcriptase Polymerase Chain Reaction
Chemical
Reg. No./Substance:
0/Antibodies; 0/CCAAT-Enhancer-Binding Protein-beta; 0/Indoles; 0/Mmp14 protein, mouse; 0/Protease Inhibitors; 142880-36-2/ilomastat; EC 3.4.24.-/Matrix Metalloproteinases; EC 3.4.24.-/Matrix Metalloproteinases, Membrane-Associated; EC 3.4.24.-/Metalloendopeptidases; EC 3.4.24.24/Matrix Metalloproteinase 2; EC 3.4.24.35/Matrix Metalloproteinase 9; EC 3.4.24.80/Matrix Metalloproteinase 14
Comments/Corrections

From MEDLINE®/PubMed®, a database of the U.S. National Library of Medicine


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