Document Detail


Intracellular acetyl unit transport in fungal carbon metabolism.
MedLine Citation:
PMID:  20889721     Owner:  NLM     Status:  MEDLINE    
Abstract/OtherAbstract:
Acetyl coenzyme A (acetyl-CoA) is a central metabolite in carbon and energy metabolism. Because of its amphiphilic nature and bulkiness, acetyl-CoA cannot readily traverse biological membranes. In fungi, two systems for acetyl unit transport have been identified: a shuttle dependent on the carrier carnitine and a (peroxisomal) citrate synthase-dependent pathway. In the carnitine-dependent pathway, carnitine acetyltransferases exchange the CoA group of acetyl-CoA for carnitine, thereby forming acetyl-carnitine, which can be transported between subcellular compartments. Citrate synthase catalyzes the condensation of oxaloacetate and acetyl-CoA to form citrate that can be transported over the membrane. Since essential metabolic pathways such as fatty acid β-oxidation, the tricarboxylic acid (TCA) cycle, and the glyoxylate cycle are physically separated into different organelles, shuttling of acetyl units is essential for growth of fungal species on various carbon sources such as fatty acids, ethanol, acetate, or citrate. In this review we summarize the current knowledge on the different systems of acetyl transport that are operational during alternative carbon metabolism, with special focus on two fungal species: Saccharomyces cerevisiae and Candida albicans.
Authors:
Karin Strijbis; Ben Distel
Publication Detail:
Type:  Journal Article; Research Support, Non-U.S. Gov't; Review     Date:  2010-10-01
Journal Detail:
Title:  Eukaryotic cell     Volume:  9     ISSN:  1535-9786     ISO Abbreviation:  Eukaryotic Cell     Publication Date:  2010 Dec 
Date Detail:
Created Date:  2010-12-03     Completed Date:  2011-03-10     Revised Date:  2011-07-28    
Medline Journal Info:
Nlm Unique ID:  101130731     Medline TA:  Eukaryot Cell     Country:  United States    
Other Details:
Languages:  eng     Pagination:  1809-15     Citation Subset:  IM    
Affiliation:
Dept. of Medical Biochemistry, Academic Medical Center, Meibergdreef 15, 1105 AZ Amsterdam, Netherlands. b.distel@amc.uva.nl
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MeSH Terms
Descriptor/Qualifier:
Acetyl Coenzyme A / metabolism
Acetyltransferases / genetics,  metabolism*
Candida albicans / enzymology*,  genetics,  metabolism
Carbon / metabolism*
Fungal Proteins / genetics,  metabolism*
Saccharomyces cerevisiae / enzymology*,  genetics,  metabolism
Chemical
Reg. No./Substance:
0/Fungal Proteins; 72-89-9/Acetyl Coenzyme A; 7440-44-0/Carbon; EC 2.3.1.-/Acetyltransferases
Comments/Corrections

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