Document Detail


In vitro assembly of the mouse U14 snoRNP core complex and identification of a 65-kDa box C/D-binding protein.
MedLine Citation:
PMID:  9582099     Owner:  NLM     Status:  MEDLINE    
Abstract/OtherAbstract:
The eukaryotic nucleolus contains a diverse population of small nucleolar RNAs (snoRNAs) that have been categorized into two major families based on evolutionarily conserved sequence elements. U14 snoRNA is a member of the larger, box C/D snoRNA family and possesses nucleotide box C and D consensus sequences. In previous studies, we have defined a U14 box C/D core motif that is essential for intronic U14 snoRNA processing. These studies also revealed that nuclear proteins that recognize boxes C/D are required. We have now established an in vitro U14 snoRNP assembly system to characterize protein binding. Electrophoretic mobility-shift analysis demonstrated that all the sequences and structures of the box C/D core motif required for U14 processing are also necessary for protein binding and snoRNP assembly. These required elements include a base paired 5',3' terminal stem and the phylogenetically conserved nucleotides of boxes C and D. The ability of other box C/D snoRNAs to compete for protein binding demonstrated that the box C/D core motif-binding proteins are common to this family of snoRNAs. UV crosslinking of nuclear proteins bound to the U14 core motif identified a 65-kDa mouse snoRNP protein that requires boxes C and D for binding. Two additional core motif proteins of 55 and 50 kDa were also identified by biochemical fractionation of the in vitro-assembled U14 snoRNP complex. Thus, the U14 snoRNP core complex is a multiprotein particle whose assembly requires nucleotide boxes C and D.
Authors:
N J Watkins; D R Newman; J F Kuhn; E S Maxwell
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Publication Detail:
Type:  Journal Article; Research Support, U.S. Gov't, Non-P.H.S.    
Journal Detail:
Title:  RNA (New York, N.Y.)     Volume:  4     ISSN:  1355-8382     ISO Abbreviation:  RNA     Publication Date:  1998 May 
Date Detail:
Created Date:  1998-05-27     Completed Date:  1998-05-27     Revised Date:  2009-11-18    
Medline Journal Info:
Nlm Unique ID:  9509184     Medline TA:  RNA     Country:  UNITED STATES    
Other Details:
Languages:  eng     Pagination:  582-93     Citation Subset:  IM    
Affiliation:
Department of Biochemistry, North Carolina State University, Raleigh 27695-7622, USA.
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MeSH Terms
Descriptor/Qualifier:
Animals
Ascitic Fluid / cytology
Cell Extracts
Cross-Linking Reagents
Hela Cells
Humans
Mice
Molecular Weight
Nucleic Acid Conformation
Oocytes
Protein Binding
RNA, Small Nuclear / chemistry,  genetics,  metabolism*
RNA-Binding Proteins / chemistry,  isolation & purification,  metabolism*
Ribonucleoproteins, Small Nuclear / biosynthesis*,  chemistry
Ultraviolet Rays
Xenopus
Chemical
Reg. No./Substance:
0/Cell Extracts; 0/Cross-Linking Reagents; 0/RNA, Small Nuclear; 0/RNA-Binding Proteins; 0/Ribonucleoproteins, Small Nuclear
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