Document Detail


In chemotaxing fibroblasts, both high-fidelity and weakly biased cell movements track the localization of PI3K signaling.
MedLine Citation:
PMID:  21504725     Owner:  NLM     Status:  MEDLINE    
Abstract/OtherAbstract:
Cell movement biased by a chemical gradient, or chemotaxis, coordinates the recruitment of cells and collective migration of cell populations. During wound healing, chemotaxis of fibroblasts is stimulated by platelet-derived growth factor (PDGF) and certain other chemoattractants. Whereas the immediate PDGF gradient sensing response has been characterized previously at the level of phosphoinositide 3-kinase (PI3K) signaling, the sensitivity of the response at the level of cell migration bias has not yet been studied quantitatively. In this work, we used live-cell total internal reflection fluorescence microscopy to monitor PI3K signaling dynamics and cell movements for extended periods. We show that persistent and properly aligned (i.e., high-fidelity) fibroblast migration does indeed correlate with polarized PI3K signaling; accordingly, this behavior is seen only under conditions of high gradient steepness (>10% across a typical cell length of 50 μm) and a certain range of PDGF concentrations. Under suboptimal conditions, cells execute a random or biased random walk, but nonetheless move in a predictable fashion according to the changing pattern of PI3K signaling. Inhibition of PI3K during chemotaxis is accompanied by loss of both cell-substratum contact and morphological polarity, but after a recovery period, PI3K-inhibited fibroblasts often regain the ability to orient toward the PDGF gradient.
Authors:
Adam T Melvin; Erik S Welf; Yana Wang; Darrell J Irvine; Jason M Haugh
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Publication Detail:
Type:  Journal Article; Research Support, N.I.H., Extramural; Research Support, Non-U.S. Gov't; Research Support, U.S. Gov't, Non-P.H.S.    
Journal Detail:
Title:  Biophysical journal     Volume:  100     ISSN:  1542-0086     ISO Abbreviation:  Biophys. J.     Publication Date:  2011 Apr 
Date Detail:
Created Date:  2011-04-20     Completed Date:  2011-08-11     Revised Date:  2012-05-04    
Medline Journal Info:
Nlm Unique ID:  0370626     Medline TA:  Biophys J     Country:  United States    
Other Details:
Languages:  eng     Pagination:  1893-901     Citation Subset:  IM    
Copyright Information:
Copyright © 2011 Biophysical Society. Published by Elsevier Inc. All rights reserved.
Affiliation:
Department of Chemical and Biomolecular Engineering, North Carolina State University, Raleigh, North Carolina, USA.
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MeSH Terms
Descriptor/Qualifier:
Alginates / chemistry
Animals
Cell Surface Extensions / drug effects,  metabolism
Chemotaxis* / drug effects
Dose-Response Relationship, Drug
Fibroblasts / cytology*,  drug effects,  enzymology*
Glucuronic Acid / chemistry
Hexuronic Acids / chemistry
Mice
Microspheres
NIH 3T3 Cells
Phosphatidylinositol 3-Kinases / metabolism*
Platelet-Derived Growth Factor / chemistry,  pharmacology
Receptors, Platelet-Derived Growth Factor / metabolism
Signal Transduction* / drug effects
Stochastic Processes
Grant Support
ID/Acronym/Agency:
EB007280/EB/NIBIB NIH HHS; GM074711/GM/NIGMS NIH HHS; R21 GM074711/GM/NIGMS NIH HHS; R21 GM074711-02/GM/NIGMS NIH HHS; //Howard Hughes Medical Institute
Chemical
Reg. No./Substance:
0/Alginates; 0/Hexuronic Acids; 0/Platelet-Derived Growth Factor; 576-37-4/Glucuronic Acid; 9005-32-7/alginic acid; EC 2.7.1.-/Phosphatidylinositol 3-Kinases; EC 2.7.10.1/Receptors, Platelet-Derived Growth Factor

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