Document Detail


Glutamate availability is important in intramuscular amino acid metabolism and TCA cycle intermediates but does not affect peak oxidative metabolism.
MedLine Citation:
PMID:  18511521     Owner:  NLM     Status:  MEDLINE    
Abstract/OtherAbstract:
Muscle glutamate is central to reactions producing 2-oxoglutarate, a tricarboxylic acid (TCA) cycle intermediate that essentially expands the TCA cycle intermediate pool during exercise. Paradoxically, muscle glutamate drops approximately 40-80% with the onset of exercise and 2-oxoglutarate declines in early exercise. To investigate the physiological relationship between glutamate, oxidative metabolism, and TCA cycle intermediates (i.e., fumarate, malate, 2-oxoglutarate), healthy subjects trained (T) the quadriceps of one thigh on the single-legged knee extensor ergometer (1 h/day at 70% maximum workload for 5 days/wk), while their contralateral quadriceps remained untrained (UT). After 5 wk of training, peak oxygen consumption (VO2peak) in the T thigh was greater than that in the UT thigh (P<0.05); VO2peak was not different between the T and UT thighs with glutamate infusion. Peak exercise under control conditions revealed a greater glutamate uptake in the T thigh compared with rest (7.3+/-3.7 vs. 1.0+/-0.1 micromol.min(-1).kg wet wt(-1), P<0.05) without increase in TCA cycle intermediates. In the UT thigh, peak exercise (vs. rest) induced an increase in fumarate (0.33+/-0.07 vs. 0.02+/-0.01 mmol/kg dry wt (dw), P<0.05) and malate (2.2+/-0.4 vs. 0.5+/-0.03 mmol/kg dw, P<0.05) and a decrease in 2-oxoglutarate (12.2+/-1.6 vs. 32.4+/-6.8 micromol/kg dw, P<0.05). Overall, glutamate infusion increased arterial glutamate (P<0.05) and maintained this increase. Glutamate infusion coincided with elevated fumarate and malate (P<0.05) and decreased 2-oxoglutarate (P<0.05) at peak exercise relative to rest in the T thigh; there were no further changes in the UT thigh. Although glutamate may have a role in the expansion of the TCA cycle, glutamate and TCA cycle intermediates do not directly affect VO2peak in either trained or untrained muscle.
Authors:
M Mourtzakis; T E Graham; J González-Alonso; B Saltin
Publication Detail:
Type:  Journal Article; Research Support, Non-U.S. Gov't     Date:  2008-05-29
Journal Detail:
Title:  Journal of applied physiology (Bethesda, Md. : 1985)     Volume:  105     ISSN:  8750-7587     ISO Abbreviation:  J. Appl. Physiol.     Publication Date:  2008 Aug 
Date Detail:
Created Date:  2008-08-05     Completed Date:  2008-10-10     Revised Date:  -    
Medline Journal Info:
Nlm Unique ID:  8502536     Medline TA:  J Appl Physiol     Country:  United States    
Other Details:
Languages:  eng     Pagination:  547-54     Citation Subset:  IM    
Affiliation:
Human Health and Nutritional Sciences, University of Guelph, Guelph, Ontario, Canada. mmourtza@healthy.uwaterloo.ca
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MeSH Terms
Descriptor/Qualifier:
Adult
Alanine Transaminase / metabolism
Amino Acids / metabolism*
Anaerobic Threshold / physiology
Blood Glucose / metabolism
Carbon Dioxide / blood
Citric Acid Cycle / physiology*
Glucagon / blood
Glutamic Acid / metabolism*
Humans
Insulin / blood
Leg / physiology
Male
Muscle, Skeletal / metabolism*
Nitrogen / metabolism
Organ Size / physiology
Oxidation-Reduction
Oxygen / blood
Oxygen Consumption / physiology
Physical Fitness / physiology
Chemical
Reg. No./Substance:
0/Amino Acids; 0/Blood Glucose; 11061-68-0/Insulin; 124-38-9/Carbon Dioxide; 56-86-0/Glutamic Acid; 7727-37-9/Nitrogen; 7782-44-7/Oxygen; 9007-92-5/Glucagon; EC 2.6.1.2/Alanine Transaminase

From MEDLINE®/PubMed®, a database of the U.S. National Library of Medicine


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