Document Detail


Glut4 is upregulated despite decreased insulin signaling during prolonged fasting in northern elephant seal pups.
MedLine Citation:
PMID:  20980624     Owner:  NLM     Status:  MEDLINE    
Abstract/OtherAbstract:
Postprandial cellular glucose uptake is dependent on an insulin-signaling cascade in muscle and adipose tissue, resulting in the translocation of the insulin-dependent glucose transporter 4 (Glut4) into the plasma membrane. Additionally, extended food deprivation is characterized by suppressed insulin signaling and decreased Glut4 expression. Northern elephant seals are adapted to prolonged fasts characterized by high levels of plasma glucose. To address the hypothesis that the fasting-induced decrease in insulin is associated with reduced insulin signaling in prolonged fasted seals, we compared the adipose protein levels of the cellular insulin-signaling pathway, Glut4 and plasma glucose, insulin, cortisol, and adiponectin concentrations between Early (n = 9; 2-3 wks postweaning) and Late (n = 8; 6-8 wks postweaning) fasted seals. Plasma adiponectin (230 ± 13 vs. 177 ± 11 ng/ml), insulin (2.7 ± 0.4 vs. 1.0 ± 0.1 μU/ml), and glucose (9.8 ± 0.5 vs. 8.0 ± 0.3 mM) decreased, while cortisol (124 ± 6 vs. 257 ± 30 nM) doubled with fasting. Glut4 increased (31%) with fasting despite the significant decreases in the cellular content of phosphatidylinositol 3-kinase as well as phosphorylated insulin receptor, insulin receptor substrate-1, and Akt2. Increased Glut4 may have contributed to the decrease in plasma glucose, but the decrease in insulin and insulin signaling suggests that Glut4 is not insulin-dependent in adipose tissue during prolonged fasting in elephant seals. The reduction of plasma glucose independent of insulin may make these animals an ideal model for the study of insulin resistance.
Authors:
Jose A Viscarra; José Pablo Vázquez-Medina; Daniel E Crocker; Rudy M Ortiz
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Publication Detail:
Type:  Journal Article; Research Support, N.I.H., Extramural; Research Support, Non-U.S. Gov't     Date:  2010-10-27
Journal Detail:
Title:  American journal of physiology. Regulatory, integrative and comparative physiology     Volume:  300     ISSN:  1522-1490     ISO Abbreviation:  Am. J. Physiol. Regul. Integr. Comp. Physiol.     Publication Date:  2011 Jan 
Date Detail:
Created Date:  2010-12-24     Completed Date:  2011-01-20     Revised Date:  2012-01-02    
Medline Journal Info:
Nlm Unique ID:  100901230     Medline TA:  Am J Physiol Regul Integr Comp Physiol     Country:  United States    
Other Details:
Languages:  eng     Pagination:  R150-4     Citation Subset:  IM    
Affiliation:
University of California, Merced, 95343, USA. jviscarra@ucmerced.edu
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MeSH Terms
Descriptor/Qualifier:
Adiponectin / blood
Adipose Tissue / metabolism
Animals
Animals, Newborn / metabolism
Blood Glucose / metabolism
Fasting / metabolism*
Glucose Transporter Type 4 / metabolism*
Hydrocortisone / blood
Insulin / metabolism*
Insulin Resistance / physiology
Models, Animal
Muscle, Skeletal / metabolism
Seals, Earless / metabolism*
Signal Transduction / physiology*
Up-Regulation / physiology*
Grant Support
ID/Acronym/Agency:
HL-091767/HL/NHLBI NIH HHS; R01 HL091767-04/HL/NHLBI NIH HHS
Chemical
Reg. No./Substance:
0/Adiponectin; 0/Blood Glucose; 0/Glucose Transporter Type 4; 0/Insulin; 50-23-7/Hydrocortisone

From MEDLINE®/PubMed®, a database of the U.S. National Library of Medicine


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