Document Detail


Gamma-interferon-inducible lysosomal thiol reductase (GILT). Maturation, activity, and mechanism of action.
MedLine Citation:
PMID:  10852914     Owner:  NLM     Status:  MEDLINE    
Abstract/OtherAbstract:
We recently identified a gamma-interferon-inducible lysosomal thiol reductase (GILT), constitutively expressed in antigen-presenting cells, that catalyzes disulfide bond reduction both in vitro and in vivo and is optimally active at acidic pH. GILT is synthesized as a 35-kDa precursor, and following delivery to major histocompatibility complex (MHC) class II-containing compartments (MIICs), is processed to the mature 30-kDa form via cleavage of N- and C-terminal propeptides. The generation of MHC class II epitopes requires both protein denaturation and reduction of intra- and inter-chain disulfide bonds prior to proteolysis. GILT may be important in disulfide bond reduction of proteins delivered to MIICs and consequently in antigen processing. In this report we show that, like its mature form, precursor GILT reduces disulfide bonds with an acidic pH optimum, suggesting that it may also be involved in disulfide bond reduction in the endocytic pathway. We also show that processing of precursor GILT can be mediated by multiple lysosomal proteases and provide evidence that the mechanism of action of GILT resembles that of other thiol oxidoreductases.
Authors:
U T Phan; B Arunachalam; P Cresswell
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Publication Detail:
Type:  Journal Article; Research Support, Non-U.S. Gov't; Research Support, U.S. Gov't, P.H.S.    
Journal Detail:
Title:  The Journal of biological chemistry     Volume:  275     ISSN:  0021-9258     ISO Abbreviation:  J. Biol. Chem.     Publication Date:  2000 Aug 
Date Detail:
Created Date:  2000-09-25     Completed Date:  2000-09-25     Revised Date:  2007-11-14    
Medline Journal Info:
Nlm Unique ID:  2985121R     Medline TA:  J Biol Chem     Country:  UNITED STATES    
Other Details:
Languages:  eng     Pagination:  25907-14     Citation Subset:  IM    
Affiliation:
Section of Immunobiology, Howard Hughes Medical Institute, Yale University School of Medicine, New Haven, Connecticut 06510, USA.
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MeSH Terms
Descriptor/Qualifier:
Animals
Antigen-Presenting Cells / enzymology*
COS Cells
Catalysis
Cell Line
Disulfides / metabolism
Endocytosis
Enzyme Precursors / isolation & purification,  metabolism*
Histocompatibility Antigens Class II / metabolism
Humans
Hydrogen-Ion Concentration
Molecular Weight
Oxidoreductases / isolation & purification,  metabolism*
Oxidoreductases Acting on Sulfur Group Donors
Protein Denaturation
Rabbits
Grant Support
ID/Acronym/Agency:
AI23081/AI/NIAID NIH HHS
Chemical
Reg. No./Substance:
0/Disulfides; 0/Enzyme Precursors; 0/Histocompatibility Antigens Class II; EC 1.-/Oxidoreductases; EC 1.8.-/IFI30 protein, human; EC 1.8.-/Oxidoreductases Acting on Sulfur Group Donors

From MEDLINE®/PubMed®, a database of the U.S. National Library of Medicine


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