Document Detail


Functional interchangeability of rod and cone transducin alpha-subunits.
MedLine Citation:
PMID:  19815523     Owner:  NLM     Status:  MEDLINE    
Abstract/OtherAbstract:
Rod and cone photoreceptors use similar but distinct sets of phototransduction proteins to achieve different functional properties, suitable for their role as dim and bright light receptors, respectively. For example, rod and cone visual pigments couple to distinct variants of the heterotrimeric G protein transducin. However, the role of the structural differences between rod and cone transducin alpha subunits (Talpha) in determining the functional differences between rods and cones is unknown. To address this question, we studied the translocation and signaling properties of rod Talpha expressed in cones and cone Talpha expressed in rods in three mouse strains: rod Talpha knockout, cone Talpha GNAT2(cpfl3) mutant, and rod and cone Talpha double mutant rd17 mouse. Surprisingly, although the rod/cone Talpha are only 79% identical, exogenously expressed rod or cone Talpha localized and translocated identically to endogenous Talpha in each photoreceptor type. Moreover, exogenously expressed rod or cone Talpha rescued electroretinogram responses (ERGs) in mice lacking functional cone or rod Talpha, respectively. Ex vivo transretinal ERG and single-cell recordings from rd17 retinas treated with rod or cone Talpha showed comparable rod sensitivity and response kinetics. These results demonstrate that cone Talpha forms a functional heterotrimeric G protein complex in rods and that rod and cone Talpha couple equally well to the rod phototransduction cascade. Thus, rod and cone transducin alpha-subunits are functionally interchangeable and their signaling properties do not contribute to the intrinsic light sensitivity differences between rods and cones. Additionally, the technology used here could be adapted for any such homologue swap desired.
Authors:
Wen-Tao Deng; Keisuke Sakurai; Jianwen Liu; Astra Dinculescu; Jie Li; Jijing Pang; Seok-Hong Min; Vince A Chiodo; Sanford L Boye; Bo Chang; Vladimir J Kefalov; William W Hauswirth
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Publication Detail:
Type:  Journal Article; Research Support, N.I.H., Extramural; Research Support, Non-U.S. Gov't     Date:  2009-10-06
Journal Detail:
Title:  Proceedings of the National Academy of Sciences of the United States of America     Volume:  106     ISSN:  1091-6490     ISO Abbreviation:  Proc. Natl. Acad. Sci. U.S.A.     Publication Date:  2009 Oct 
Date Detail:
Created Date:  2009-10-21     Completed Date:  2009-11-23     Revised Date:  2010-09-28    
Medline Journal Info:
Nlm Unique ID:  7505876     Medline TA:  Proc Natl Acad Sci U S A     Country:  United States    
Other Details:
Languages:  eng     Pagination:  17681-6     Citation Subset:  IM    
Affiliation:
Department of Ophthalmology, University of Florida, Gainesville, FL 32610, USA. wdeng@ufl.edu
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MeSH Terms
Descriptor/Qualifier:
Animals
Electroretinography
Evoked Potentials, Visual
Eye Proteins / chemistry*,  genetics,  physiology*
Heterotrimeric GTP-Binding Proteins / chemistry*,  deficiency,  genetics,  physiology*
Mice
Mice, Knockout
Mice, Mutant Strains
Photic Stimulation
Protein Subunits
Recombinant Proteins / chemistry,  genetics,  metabolism
Retinal Cone Photoreceptor Cells / physiology*
Retinal Rod Photoreceptor Cells / physiology*
Grant Support
ID/Acronym/Agency:
EY02687/EY/NEI NIH HHS; EY08571/EY/NEI NIH HHS; EY11123/EY/NEI NIH HHS; NS36302/NS/NINDS NIH HHS
Chemical
Reg. No./Substance:
0/Eye Proteins; 0/Gnat2 protein, mouse; 0/Protein Subunits; 0/Recombinant Proteins; EC 3.6.5.1/Heterotrimeric GTP-Binding Proteins
Comments/Corrections

From MEDLINE®/PubMed®, a database of the U.S. National Library of Medicine


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