Document Detail


Functional interaction of the circadian clock and UV RESISTANCE LOCUS 8-controlled UV-B signaling pathways in Arabidopsis thaliana.
MedLine Citation:
PMID:  21395889     Owner:  NLM     Status:  Publisher    
Abstract/OtherAbstract:
Circadian clocks regulate many molecular and physiological processes in Arabidopsis (Arabidopsis thaliana) with a 24 h period, allowing the timing of these processes to the most appropriate time of the day. The accuracy of timing relies on the synchrony of the clock and the environmental day/night cycle. Visible light is the most potent signal for such synchronization but light-induced responses are also rhythmically attenuated (gated) by the clock. Here we report a similar mutual interaction of the circadian clock and non-damaging photomorphogenic UV-B light. We show that low intensity UV-B radiation acts as entraining signal for the clock. UV RESISTANCE LOCUS 8 (UVR8) and CONSTITUTIVELY PHOTOMORPHOGENIC 1 (COP1) are required, but ELONGATED HYPOCOTYL 5 (HY5) and HY5 HOMOLOG (HYH), are dispensable for this process. UV-B responsiveness of clock gene expression suggests that photomorphogenic UV-B entrains the plant clock through transcriptional activation. We also demonstrate that UV-B induction of gene expression under these conditions is gated by the clock in a HY5/HYH-independent manner. The arrhythmic early flowering 3-4 mutant showed non-gated, high level gene induction by UV-B, yet displayed no increased tolerance to UV-B stress. Thus, the temporal restriction of UV-B responsiveness by the circadian clock can be considered as saving resources during acclimation without losing fitness.
Authors:
Balázs Fehér; László Kozma-Bognár; Eva Kevei; Anita Hajdu; Melanie Binkert; Seth Jon Davis; Eberhard Schäfer; Roman Ulm; Ferenc Nagy
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Publication Detail:
Type:  JOURNAL ARTICLE     Date:  2011-3-14
Journal Detail:
Title:  The Plant journal : for cell and molecular biology     Volume:  -     ISSN:  1365-313X     ISO Abbreviation:  -     Publication Date:  2011 Mar 
Date Detail:
Created Date:  2011-3-14     Completed Date:  -     Revised Date:  -    
Medline Journal Info:
Nlm Unique ID:  9207397     Medline TA:  Plant J     Country:  -    
Other Details:
Languages:  ENG     Pagination:  -     Citation Subset:  -    
Copyright Information:
Copyright © 2011 Blackwell Publishing Ltd and the Society for Experimental Biology.
Affiliation:
Institute of Plant Biology, Biological Research Centre of the Hungarian Academy of Sciences, H-6726 Szeged, Hungary Faculty of Biology, Institute of Biology II/Botany, University of Freiburg, D-79104 Freiburg, Germany Present address: Department of Plant Biology, University of Geneva, Sciences III, CH-1211 Geneva 4, Switzerland Department of Plant Developmental Biology, Max Planck Institute for Plant Breeding Research, D-50829, Cologne, Germany Centre for Biological Signalling Studies (BIOSS), University of Freiburg, D-79104 Freiburg, Germany School of Biological Sciences, University of Edinburgh, EH9 3JR Edinburgh, United Kingdom.
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