Document Detail


Follicular exclusion and rapid elimination of hen egg lysozyme autoantigen-binding B cells are dependent on competitor B cells, but not on T cells.
MedLine Citation:
PMID:  9886397     Owner:  NLM     Status:  MEDLINE    
Abstract/OtherAbstract:
In mice with a diverse B cell repertoire, hen egg lysozyme (HEL) autoantigen-binding B cells are excluded from follicles and eliminated in 3 days. To explore the roles of competitor B cells and of T cells in this mechanism of self-tolerance, HEL-specific B cells were transferred into mice containing HEL and deficient in endogenous B cells (muMT), T cells (TCR-/-), or B and T cells (RAG1-/-). Previous studies suggested a dual requirement for B cell receptor (BCR) engagement and competition in HEL autoantigen-binding B cell elimination, but interpretation of these experiments has been confounded by the possible failure to independently regulate autoantigen concentration and competitor B cell frequency. In experiments in this study, we have fixed one variable, HEL concentration, while varying the second, the presence or absence of other B cells. By this approach, we find that follicular exclusion and rapid elimination of autoreactive B cells require BCR engagement plus competition with other B cells, rather than BCR engagement alone. We also find, by transfers into T cell-deficient mice, that T cells are not required for this peripheral tolerance mechanism. Unexpectedly, in mice lacking both T cells and competitor B cells (RAG1-/-), transferred HEL-binding cells survive less well than in mice just lacking competitor B cells. These results suggest T cells can enhance autoreactive B cell survival. Enhanced survival of autoreactive B cells, due to the presence of T cells and the lack of competitor B cells, might contribute to the elevated frequency of autoimmunity in B cell-deficient individuals.
Authors:
K N Schmidt; J G Cyster
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Publication Detail:
Type:  Journal Article; Research Support, Non-U.S. Gov't; Research Support, U.S. Gov't, P.H.S.    
Journal Detail:
Title:  Journal of immunology (Baltimore, Md. : 1950)     Volume:  162     ISSN:  0022-1767     ISO Abbreviation:  J. Immunol.     Publication Date:  1999 Jan 
Date Detail:
Created Date:  1999-01-21     Completed Date:  1999-01-21     Revised Date:  2007-11-14    
Medline Journal Info:
Nlm Unique ID:  2985117R     Medline TA:  J Immunol     Country:  UNITED STATES    
Other Details:
Languages:  eng     Pagination:  284-91     Citation Subset:  AIM; IM    
Affiliation:
Department of Microbiology and Immunology, University of California, San Francisco 94143, USA.
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MeSH Terms
Descriptor/Qualifier:
Animals
Autoantigens / metabolism*
B-Lymphocyte Subsets / immunology*,  metabolism,  pathology
Binding, Competitive / immunology
Cell Movement / immunology*
Cell Survival / immunology
Chickens
Lymphoid Tissue / immunology*,  metabolism,  pathology
Lymphopenia / genetics,  immunology,  pathology
Mice
Mice, Inbred C57BL
Mice, Knockout
Mice, Transgenic
Muramidase / immunology*,  metabolism
T-Lymphocyte Subsets / immunology*,  metabolism
Grant Support
ID/Acronym/Agency:
AI 40098/AI/NIAID NIH HHS
Chemical
Reg. No./Substance:
0/Autoantigens; EC 3.2.1.17/Muramidase

From MEDLINE®/PubMed®, a database of the U.S. National Library of Medicine


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